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manyray aster

arrow-leaf American-aster, white arrowleaf aster, white arrowleaf or arrowleaf aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, 40–120(–200) cm, cespitose; short-rhizomatous or with stout, branched caudices.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–5+, erect (straight to slightly flexuous, brittle, sometimes stout), proximally glabrous or glabrate, distally sparsely pilose.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

thin, margins usually shallowly, sometimes deeply serrate, rarely entire (distal), (piloso- to strigoso-)ciliate to scabrous (distal), apices mucronate, abaxial faces moderately pilose, ± densely so on midveins, adaxial glabrate to sparsely pilose or scabrous;

basal usually withering by flowering, sometimes persistent, long-petiolate (5–15 cm, petioles not or narrowly winged, sheathing), blades ovate to lance-ovate or lanceolate, 25–120 × 15–50 mm, bases usually shallowly, sometimes deeply, cordate to truncate or rounded, apices acute, sometimes rounded;

proximal cauline sometimes deciduous at flowering, winged-petiolate (petioles distally usually broader-winged, clasping), blades ovate to lanceolate, 50–120 × 20–50 mm, gradually reduced distally, bases usually truncate or rounded to attenuate, sometimes cordate, apices usually acute or acuminate, sometimes nearly caudate;

distal ± shortly winged-petiolate or sessile, blades lance-ovate or lanceolate to linear-lanceolate, 10–100 × 1–25 mm, reduced distally, bases attenuate to cuneate, margins entire to subentire, scabrous, apices acuminate to subcaudate.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.3–2 cm, sparsely pilosulous to glabrate, bracts many, ascending, linear to subulate, 2–4 mm, grading into phyllaries.

Involucres

campanulate, 5–10 mm.

cylindric, (4–)4.5–6(–7) mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

8–15(–20);

corollas mostly white to pale pink or lilac or light blue, laminae 4.5–8(–10) × 0.8–1.5 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

(8–)10–15(–20);

corollas whitish to cream becoming pink, (3.5–)4–5 mm, tubes shorter than funnelform throats (with hairs), lobes lanceolate, 0.4–0.7 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 4–6 series, appressed or recurved-spreading, subulate (outer) to linear-lanceolate (inner), unequal, bases indurate 1/3–1/2+, margins scarious, hyaline, erose, sparsely ciliolate (mostly distally), green zones linear-lanceolate (outer) to linear (inner), apices long-acuminate to caudate, recurved, hyaline-mucronate to aristate, tips often purple, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

(crowded) in dense, elongate, paniculiform arrays, branches stiffly ascending to erect (sometimes arched when long, peduncles then often secund), usually racemiform, sometimes paniculiform, leafy.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

tan to brown or dull purple (nerves stramineous), obovoid, ± compressed, 1.8–2.5 mm, 4–6-nerved, faces glabrous or glabrate or very sparsely strigillose;

pappi white or rose-tinged, 3–4 mm.

2n

= 16.

= 16.

Symphyotrichum anomalum

Symphyotrichum urophyllum

Phenology Flowering late Aug–Oct. Flowering Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams Open, dry or mesic, sandy or loamy, sometimes rocky soils, limestone pavements, glades, ravines, open oak, sassafras, aspen, or pine woods, edges of woods, moist or dry, rocky bluffs, savanna, wooded banks, stabilized dunes, fields, roadsides, hedgerows, railroads
Elevation 50–500 m (200–1600 ft) 40–300+ m (100–1000+ ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NJ; NY; OH; OK; PA; SC; TN; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Aster sagittifolius Wedemeyer ex Willdenow has been misapplied to this taxon; our interpretation of the type of this entity concords with that of A. G. Jones (1980), i.e., that it is conspecific with the type of Symphyotrichum cordifolium or a garden hybrid involving the latter. Aster sagittifolius forma hirtellus (Lindley) Shinners is not recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 504. FNA vol. 20, p. 506.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Synonyms Aster anomalus Aster urophyllus, Aster sagittifolius var. dissitiflorus
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Lindley ex de Candolle) G. L. Nesom: Phytologia 77: 294. (1995)
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