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manyray aster

aster poilu, awl American-aster, frost aster, hairy aster, hairy white Oldfield aster, Oldfield or frost weed or white Oldfield or hairy or hairy white Oldfield aster, white heath aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials cespitose, (5–)20–120(–150+) cm; with stout, branched caudices, sometimes long-rhizomatous.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–5+, ascending to erect (straight, stout), glabrate to hirsuto-hispid or pilose, sometimes glabrous or hairy in lines (var. pringlei).

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

thin, margins ciliate, apices acute to acuminate, mucronate, faces glabrous or ± hirsute (particularly on abaxial midveins);

basal withering by flowering (new vernal rosettes developing at flowering), petiolate to subpetiolate (petioles winged, ciliate, bases sheathing), blades oblanceolate or obovate to spatulate, 10–60 × 5–15 mm, bases attenuate, margins sparsely crenate-serrate (mostly apically), apices obtuse to rounded;

proximal cauline usually deciduous at flowering (often with axillary clusters of small leaves), petiolate or subpetiolate to subsessile (petioles narrowly to broadly winged, clasping), blades elliptic-oblanceolate or -oblong to linear-lanceolate or -oblanceolate, 40–102 × 5–25 mm, progressively reduced distally, bases attenuate to cuneate, ± clasping, margins entire to serrate, softly ciliate, apices attenuate, hyaline-spinulose;

distal sessile, blades lance-oblong to linear-lanceolate to linear or linear-oblanceolate or linear-subulate, 10–100 × 1–8 mm, progressively reduced distally, branch leaves abruptly smaller, bases cuneate, margins entire or serrulate.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

5–30(–50) mm, progressively shorter distally, glabrous or densely hispid, bracts 7–25+, appressed to ascending, sometimes ± spreading, linear to subulate, glabrate, ciliate or not, mucronate to apiculate, ± grading into phyllaries.

Involucres

campanulate, 5–10 mm.

campanulate to cylindro-campanulate, (2.5–)3.5–5.1(–6.5) mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

(10–)16–28(–38);

corollas usually white, rarely pinkish or bluish, laminae (4–)5.4–7.5(–11) × (0.4–)0.8–1.3(–1.7) mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

(13–)17–39(–67);

corollas light yellow becoming reddish purple or brown, (2.5–)3–4.1(–5.5) mm, tubes much shorter than narrowly funnelform throats, lobes lanceolate, (0.4–)0.6–0.8(–1) mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 4–6 series, appressed or slightly spreading, oblong-lanceolate (outer) to linear (innermost) , unequal (rarely subequal), bases indurate 1/2–3/5, margins hyaline, scarious, erose, ± ciliolate distally, green zones lanceolate to lance-rhombic, apices acute or acuminate, involute, spinulose, faces glabrous or sparsely hirsutulous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in open, leafy, often pyramidal, paniculiform arrays, branches divaricate or branches 10 cm or less ascending, often arched and secund, sometimes racemiform.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

whitish or gray, oblong-obovoid, sometimes ± compressed, 1–1.5 × 0.5–0.7 mm, 4–6-nerved, faces sparsely strigillose;

pappi white, 3.5–4 mm.

2n

= 16.

Symphyotrichum anomalum

Symphyotrichum pilosum

Phenology Flowering late Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams
Elevation 50–500 m (200–1600 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; BC; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The ecology and cytogeography of Symphyotrichum pilosum were summarized by J. G. Chmielewski and J. C. Semple (2001b). The species is often confused with S. ericoides, which inhabits the same kinds of open, prairie-like habitats; the latter can be distinguished by its smaller heads and spine-tipped phyllary apices. Because of that, the name Aster ericoides has been misapplied often in floras and in the horticultural trade.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems sparsely to densely hirsute; leaves pilose; various substrates
var. pilosum
1. Stems glabrous; leaves glabrous or glabrate; mainly on calcareous substrates
var. pringlei
Source FNA vol. 20, p. 504. FNA vol. 20, p. 511.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. pilosum var. pilosum, S. pilosum var. pringlei
Synonyms Aster anomalus Aster pilosus, Aster ericoides var. pilosus
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Willdenow) G. L. Nesom: Phytologia 77: 289. (1995)
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