Symphyotrichum anomalum |
Symphyotrichum lateriflorum |
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manyray aster |
aster latériflore, calico American-aster, calico aster, calico or one-side or white woodland or starved aster |
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Habit | Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. | Perennials, 20–120(–150) cm, cespitose; with short, woody, branched caudices, or short-rhizomatous. |
Stems | 1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute. |
1–5+, ascending to erect (± arching, slender, brittle), ± densely pilose or villous to glabrate or glabrous (particularly proximally). |
Leaves | thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils); basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded; proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point; distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate. |
thin, membranous, margins serrate to serrulate, scabridulous, apices acute to acuminate, sometimes caudate, mucronulate, abaxial faces glabrous, midveins usually ± densely pilose to glabrate, rarely glabrous, adaxial scabrous (short-strigose) to glabrate; basal usually withering by flowering, sometimes persisting (new vernal rosettes often developing at flowering), petiolate to subpetiolate (petioles narrowly to ± broadly winged, ciliate, bases ± sheathing), blades oblanceolate, lance-ovate or ovate to spatulate or suborbiculate, 5–35 × 7–25 mm, abruptly attenuate, margins crenate-serrate, apices obtuse to rounded or acute; proximal cauline mostly withering by flowering, sessile or subpetiolate (petioles broadly winged), blades usually ovate or elliptic to elliptic-oblanceolate or lanceolate, rarely linear-lanceolate, (30–)50–100(–150) × (2–)10–20(–35) mm, greatly reduced distally, bases attenuate to cuneate; distal sessile, blades ovate, lance-ovate, lance-elliptic, or oblanceolate to lance-linear or linear, 10–150 × 1–30 mm, progressively reduced distally, branch leaves abruptly smaller, bases cuneate, margins sometimes entire. |
Peduncles | densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries. |
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Involucres | campanulate, 5–10 mm. |
cylindro-campanulate, (3.5–)4–6(–7) mm. |
Ray florets | 18–45; corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm. |
8–15(–23); corollas white, rarely pinkish or purplish, laminae (3–)4–5(–8) × 0.9–1.2 mm. |
Disc florets | 33–40+; corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm. |
8–16(–20); corollas cream to light yellow turning pink or reddish purple, (2.5–)3–5 mm, tubes shorter than funnelform-campanulate throats, lobes strongly reflexed, lanceolate, 0.9–1.7 mm. |
Phyllaries | in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous. |
in 3–4(–6) series, appressed or slightly spreading, oblong-lanceolate or -oblanceoalte (outer) to linear (inner), unequal, bases indurate 1/3–2/3, margins scarious, erose, hyaline or sometimes reddish, ± ciliolate, green zones lanceolate to ± diamond-shaped, apices (outer) acute, callus-pointed, (mid) acute to acuminate, or obtuse, sometimes purplish, abaxial faces glabrous (outer), mid sparsely puberulent. |
Heads | in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate. |
in ample, open, diffuse, ± pyramidal, paniculiform arrays, branches divaricate to long-arching or ± ascending, slender, wiry, secund; usually sessile, sometimes peduncles 0.1–1 cm+, ± pilose, bracts 1–7, linear or subulate to oblong-lanceolate, foliaceous, grading into phyllaries. |
Cypselae | deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous; pappi tawny or rose-tinged, 3.7–5.2 mm. |
gray or tan, oblong-obovoid, sometimes ± compressed, (1.3–)1.8–2.2 mm, 3–5-nerved, faces sparsely strigillose; pappi white to pinkish, 3–4 mm. |
2n | = 16. |
= 16, 32, 48. |
Symphyotrichum anomalum |
Symphyotrichum lateriflorum |
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Phenology | Flowering late Aug–Oct. | Flowering Aug–Oct. |
Habitat | Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams | Relatively shaded habitats, dry to humid soils, usually deciduous woodlands (beech–maple, oak–hickory, mixed hardwoods), sometimes conifer woods, sometimes swamp forests (conifer or dediduous), edges of woods, meadows, usually rocky or sandy stream and pond shores, roadsides |
Elevation | 50–500 m (200–1600 ft) | 0–400 m (0–1300 ft) |
Distribution |
AR; IL; KS; MO; OK
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC
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Discussion | A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum lateriflorum has been reported in British Columbia as an ephemeral that did not persist. G. L. Nesom (1994b) and J. C. Semple et al. (2002) recognized several varieties within the complex: var. lateriflorum (syn. Aster vimineus Lamarck); var. angustifolium (Wiegand) G. L. Nesom (syn. A. lateriflorus var. angustifolius Wiegand); var. flagellare (Shinners) G. L. Nesom (syn. A. lateriflorus var. flagellaris Shinners, A. lateriflorus var. indutus Shinners); var. hirsuticaule (Lindley ex de Candolle) G. L. Nesom [syn. A. hirsuticaulis Lindley ex de Candolle, A. lateriflorus var. hirsuticaulis (Lindley ex de Candolle) Porter]; var. horizontale (Desfontaines) G. L. Nesom [syn. A. horizontalis Desfontaines, A. lateriflorus var. horizontalis (Desfontaines) Farwell]; var. spatelliforme (E. S. Burgess) G. L. Nesom [syn. A. spatelliformis E. S. Burgess, A. lateriflorus var. spatelliformis (E. S. Burgess) A. G. Jones]; and var. tenuipes (Wiegand) G. L. Nesom (syn. A. lateriflorus var. tenuipes Wiegand, A. acadiensis Shinners). Much genetic and phenotypic variation is encountered within the complex; a thorough study is needed before a coherent taxonomy can be achieved. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 504. | FNA vol. 20, p. 517. |
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum |
Sibling taxa | ||
Synonyms | Aster anomalus | Solidago lateriflora, Aster lateriflorus |
Name authority | (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) | (Linnaeus) Á. Löve & D. Löve: Taxon 31: 359. (1982) |
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