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manyray aster

aster lancéolé, lance-leaf American-aster, lance-leaf aster, lance-leaf or panicle or white panicle aster, marsh aster, panicle aster, tall white aster, western willow aster, white-panicle aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, 30–150(–200) cm, colonial; long-rhizomatous (rhizomes usually white, thick, contorted).
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1, erect (straight, stout), glabrous or hairy.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

thin, scabrous, apices mucronate, abaxial faces glabrous (vein areoles indistinct, elongate), adaxial glabrous or sparsely scabrous (var. hesperium);

basal withering by flowering, petiolate to subpetiolate (petioles winged, ciliate, bases dilated, sheathing), blades elliptic-oblanceolate or obovate to suborbiculate, 10–80 × 5–20 mm, cuneate to attenuate, margins crenate-serrate, apices acute to obtuse or rounded;

proximal cauline withering by flowering, sessile or subsessile, blades lance-ovate or oblanceolate to linear-lanceolate, (40–)50–150 × (3–)10–20(–35) mm, progressively reduced distally, bases cuneate, ± decurrent, margins serrate, apices acute to acuminate;

distal sessile, blades oblanceolate to linear, 30–100(–140) mm, only slightly reduced distally, bases cuneate, margins entire.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.5–5 cm, ± pilose, bracts 1–3(–5), linear-oblanceolate to -lanceolate, foliaceous, ciliate.

Involucres

campanulate, 5–10 mm.

campanulate to cylindric, 3–8 mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

16–50;

corollas white to pinkish or pale blue-violet, laminae 3–10(–14) × 0.5–1.3 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

(13–)20–40(–52);

corollas yellow becoming purple, 2.8–5.8 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular, 0.4–1.2 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in (3–)4–6 series, appressed or slightly spreading, linear-lanceolate to linear (innermost), sometimes slightly dilated distally, ± strongly unequal to subequal (var. hesperium), bases indurate 1/4–1/2, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate to linear-lanceolate, outer sometimes foliaceous (particularly var. hesperium), apices acute to acuminate (outer) or acuminate to caudate (inner), mostly mucronulate, abaxial faces glabrous, adaxial sparsely strigillose.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in ample or diffuse to narrow, elongate, leafy, paniculiform arrays, branches ± ascending, rarely secund, branch leaves often longer than pedicels.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

gray or tan, obovoid, ± compressed, 1.5–2 mm, 4–5-nerved, faces sparsely strigillose;

pappi white to sordid or tawny, 5.5–6 mm.

2n

= 16.

Symphyotrichum anomalum

Symphyotrichum lanceolatum

Phenology Flowering late Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams
Elevation 50–500 m (200–1600 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK [Introduced in Europe]
[WildflowerSearch map]
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (5 in the flora).

Varieties of Symphyotrichum lanceolatum are sometimes grouped into two subspecies: subsp. hesperium (var. hesperium) and subsp. lanceolatum (the other four varieties). J. C. Semple and J. G. Chmielewski (1987) provided maps of the five taxa. The ranges of var. hesperium and var. lanceolatum overlap in the prairies and in the boreal zone from Alberta to the Clay Belt of northern Ontario. The ranges of the other three varieties do not overlap with that of var. hesperium, but they all overlap each other and with var. lanceolatum in the Midwest. The ecology and cytogeography of the species were summarized by Chmielewski and Semple (2001). The name Aster tradescantii has sometimes been misapplied to this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Outer phyllaries 2/3 + length of inner ones; heads usually subtended by large foliaceous bracts; cordilleras and prairies (subsp. hesperium)
var. hesperium
1. Outer phyllaries 1/3 – 2/3 length of inner ones; heads not usually subtended by large foliaceous bracts; prairies eastwards (subsp. lanceolatum)
→ 2
2. Stems stout, moderately to densely short-woolly from bases; Great Lakes region to se Manitoba
var. hirsuticaule
2. Stems stout to slender, glabrous or at most hairy in lines at bases
→ 3
3. Heads congested on lateral branches of arrays; involucres 3–4 mm; midwestern United States, s Ontario, s Quebec
var. interior
3. Heads not congested or congested only distally on lateral branches of arrays; involucres 3.6–5.6 mm (mostly longer than 4 mm)
→ 4
4. Leaves broadly oblanceolate, not much reduced in arrays; involucres usually 4–5.5 mm; rays white
var. latifolium
4. Leaves linear to oblanceolate, reduced in arrays; involucres 3.5–5(–6) mm; rays white to purplish
var. lanceolatum
Source FNA vol. 20, p. 504. FNA vol. 20.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. lanceolatum var. hesperium, S. lanceolatum var. hirsuticaule, S. lanceolatum var. interior, S. lanceolatum var. lanceolatum, S. lanceolatum var. latifolium
Synonyms Aster anomalus Aster lanceolatus
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Willdenow) G. L. Nesom: Phytologia 77: 284. (1995)
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