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manyray aster

aster lisse, Geyer's aster, smooth American-aster, smooth aster, smooth blue aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials (15–)20–70(–120) cm, cespitose; stoutly short-rhizomatous, with thick, woody caudices or a few, long rhizomes.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–5+, erect (straight, glaucous, sometimes reddish proximally), glabrous, sometimes sparsely hirsute distally.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

(glaucous) thick, firm, ± fleshy, margins crenate-serrate or -serrulate or entire, scabridulous, apices mucronulate, faces glabrous;

basal usually withering by flowering or sometimes persistent (var. purpuratum), petiolate (petioles ± winged, bases dilated, sheathing), blades spatulate or oblong to ovate or lanceolate-ovate, 30–200 × 10–25(–30) mm, bases attenuate or cuneate to rounded, margins crenate-serrate to serrulate, apices acute to obtuse or rounded;

proximal cauline often withering by flowering, petiolate or subsessile or sessile (petioles narrowly to broadly winged, clasping), blades ovate or oblong-ovate to lanceolate or linear-lanceolate, sometimes ± panduriform, (40–)80–150(–180) × (10–)20–45 mm, bases attenuate to rounded or ± shallowly auriculate-clasping, margins entire or shallowly crenate-serrate, minutely scabrous, apices acute or obtuse, callus-pointed;

distal sessile, blades lance-ovate or lanceolate to linear, 7–45 × 1–14 mm, progressively reduced distally (abruptly so in arrays), bases auriculate and ± clasping to rounded, margins entire.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.2–6+ cm, glaucous, glabrous or puberulent in lines, bracts 3–6, densely spaced, subulate or linear-lanceolate to linear, subclasping, apices sometimes purplish, grading into phyllaries.

Involucres

campanulate, 5–10 mm.

campanulate to cylindro-campanulate, (4.2–)5–7(–8) mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

(11–)13–23(–34);

corollas usually pale to dark blue or purple, seldom white, laminae (6–)7.2–11.3(–14.6) × 1.5–2.5 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

(17–)19–33(–43);

corollas yellow turning purplish red, 3.5–6.1 mm, tubes slightly shorter than funnelform throats, lobes triangular, (0.4–)0.6–1 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 4–6 series, appressed, subulate or lanceolate (outer) to oblong-lanceolate or linear-lancolate or -oblanceolate, unequal (sometimes appearing subequal), bases indurate 1/2–3/4, margins scarious, erose, hyaline, ciliolate distally, green zones mostly diamond-shaped to ± lanceolate (some inner, or most in var. geyeri), apices acute to acuminate, sometimes ± obtuse, red-mucronate or apiculate, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in broad, sometimes ± flat-topped, paniculiform arrays, branches stiffly ascending (rarely arching), leafy with small, gradually reduced branch leaves.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

deep purple to brown, oblong-obovoid, compressed, 2–3.5 mm, 4–5-nerved, faces glabrous or glabrate;

pappi tawny to red- or rose-tinged, 5–7 mm.

2n

= 16.

Symphyotrichum anomalum

Symphyotrichum laeve

Phenology Flowering late Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams
Elevation 50–500 m (200–1600 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; ON; QC; SK; YT; ne Mexico [Introduced in other areas of Mexico, Central America, Europe]
[WildflowerSearch map]
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves lanceolate to ovate, conspicuously auriculate-clasping, lengths less than 5 times widths
→ 2
1. Leaves linear-lanceolate to linear, slightly auriculate-clasping, lengths often 5 times widths
→ 3
2. Phyllaries strongly unequal, green apical zones diamond-shaped
var. laeve
2. Phyllaries unequal, green apical zones lanceolate
var. geyeri
3. Basal leaves withering by flowering, cauline linear-lanceolate
var. concinnum
3. Leaves mostly basal and proximal at flowering, mostly linear
var. purpuratum
Source FNA vol. 20, p. 504. FNA vol. 20, p. 508.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. laeve var. concinnum, S. laeve var. geyeri, S. laeve var. laeve, S. laeve var. purpuratum
Synonyms Aster anomalus Aster laevis
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Linnaeus) Á. Löve & D. Löve: Taxon 31: 359. (1982)
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