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manyray aster

glossy-leaf aster, shiny-leaf aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, 40–250 cm, colonial; long-rhizomatous.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1, erect (straight, ± thin, 2–8 mm diam at base, ± ribbed, red above each node), glabrous or glabrate (very sparsely hispidulous) proximally to ± hispidulous distally.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

(crowded, light green, shiny) firm, margins crenulate-serrate or entire, revolute, apices acute to acuminate, mucronate, abaxial faces glabrous or midveins sometimes with hairs apically, adaxial glabrous;

basal withering by flowering, subpetiolate (petioles dilated, winged, sheathing), blades spatulate to oblanceolate, 30–100+ × 3–20+ mm, bases attenuate to cuneate, margins remotely crenate-serrate to subentire, apices acute to rounded;

proximal cauline withering by flowering, sessile or subpetiolate (petioles widely winged, clasping), blades oblanceolate, 50–150 × 20–30 mm, greatly reduced distally, bases auriculate, clasping, apices acute to acuminate;

distal sessile, blades lanceolate to lance-elliptic to oblanceolate, 40–70 × 10–25 mm, little reduced distally, bases auriculate, clasping, margins entire, apices acute to acuminate.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.2–3+ cm, glabrous or pilose in lines, bracts 4–6, lanceolate-linear, often subtending heads.

Involucres

campanulate, 5–10 mm.

campanulate, 6–12 mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

20–40;

corollas usually blue to pale lavender, sometimes white, laminae 9–18 × 1.0–1.2 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

30–50;

corollas yellow or cream becoming pink or purple, (4.5–)5–6.4 mm, tubes shorter than funnelform throats, lobes triangular to lanceolate, 0.6–0.9 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 4–5(–6) series, linear-lanceolate to linear, slightly unequal, bases indurate 1/5–1/2, margins not scarious (outer) to narrowly scarious, erose, hyaline, sparsely ciliolate, green zones linear-lanceolate, outer sometimes ± foliaceous, apices acute to acuminate or long-acuminate to caudate, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in densely paniculiform arrays, branches ascending, densely leafy (branch leaves often overtopping heads).

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

purple or brown, obovoid, oblong or oblanceolate, ± falcate, ± compressed, 1.5–3 mm, 3–4-nerved, faces glabrous or sparsely strigillose;

pappi white, 5.2–8 mm.

2n

= 16.

= 16.

Symphyotrichum anomalum

Symphyotrichum firmum

Phenology Flowering late Aug–Oct. Flowering Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams Open, wet soils, spreading into mesic mineral soils, fens, marshes, wet roadsides
Elevation 50–500 m (200–1600 ft) 100–400 m (300–1300 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
GA; IA; MI; MN; MO; NE; AB; MB; ON; SK
[WildflowerSearch map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The range of this little known species is badly defined because some specimens attributed to it are in fact white-rayed, glabrate forms of Symphyotrichum puniceum. More work is needed to verify the status of this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 504. FNA vol. 20, p. 526.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Synonyms Aster anomalus Aster firmus, Aster lucidulus, Aster puniceus var. firmus
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Nees) G. L. Nesom: Phytologia 77: 282. (1995)
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