Symphyotrichum anomalum |
Symphyotrichum firmum |
|
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manyray aster |
glossy-leaf aster, shiny-leaf aster |
|
Habit | Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. | Perennials, 40–250 cm, colonial; long-rhizomatous. |
Stems | 1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute. |
1, erect (straight, ± thin, 2–8 mm diam at base, ± ribbed, red above each node), glabrous or glabrate (very sparsely hispidulous) proximally to ± hispidulous distally. |
Leaves | thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils); basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded; proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point; distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate. |
(crowded, light green, shiny) firm, margins crenulate-serrate or entire, revolute, apices acute to acuminate, mucronate, abaxial faces glabrous or midveins sometimes with hairs apically, adaxial glabrous; basal withering by flowering, subpetiolate (petioles dilated, winged, sheathing), blades spatulate to oblanceolate, 30–100+ × 3–20+ mm, bases attenuate to cuneate, margins remotely crenate-serrate to subentire, apices acute to rounded; proximal cauline withering by flowering, sessile or subpetiolate (petioles widely winged, clasping), blades oblanceolate, 50–150 × 20–30 mm, greatly reduced distally, bases auriculate, clasping, apices acute to acuminate; distal sessile, blades lanceolate to lance-elliptic to oblanceolate, 40–70 × 10–25 mm, little reduced distally, bases auriculate, clasping, margins entire, apices acute to acuminate. |
Peduncles | densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries. |
0.2–3+ cm, glabrous or pilose in lines, bracts 4–6, lanceolate-linear, often subtending heads. |
Involucres | campanulate, 5–10 mm. |
campanulate, 6–12 mm. |
Ray florets | 18–45; corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm. |
20–40; corollas usually blue to pale lavender, sometimes white, laminae 9–18 × 1.0–1.2 mm. |
Disc florets | 33–40+; corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm. |
30–50; corollas yellow or cream becoming pink or purple, (4.5–)5–6.4 mm, tubes shorter than funnelform throats, lobes triangular to lanceolate, 0.6–0.9 mm. |
Phyllaries | in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous. |
in 4–5(–6) series, linear-lanceolate to linear, slightly unequal, bases indurate 1/5–1/2, margins not scarious (outer) to narrowly scarious, erose, hyaline, sparsely ciliolate, green zones linear-lanceolate, outer sometimes ± foliaceous, apices acute to acuminate or long-acuminate to caudate, faces glabrous. |
Heads | in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate. |
in densely paniculiform arrays, branches ascending, densely leafy (branch leaves often overtopping heads). |
Cypselae | deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous; pappi tawny or rose-tinged, 3.7–5.2 mm. |
purple or brown, obovoid, oblong or oblanceolate, ± falcate, ± compressed, 1.5–3 mm, 3–4-nerved, faces glabrous or sparsely strigillose; pappi white, 5.2–8 mm. |
2n | = 16. |
= 16. |
Symphyotrichum anomalum |
Symphyotrichum firmum |
|
Phenology | Flowering late Aug–Oct. | Flowering Aug–Oct. |
Habitat | Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams | Open, wet soils, spreading into mesic mineral soils, fens, marshes, wet roadsides |
Elevation | 50–500 m (200–1600 ft) | 100–400 m (300–1300 ft) |
Distribution |
AR; IL; KS; MO; OK
|
GA; IA; MI; MN; MO; NE; AB; MB; ON; SK
|
Discussion | A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The range of this little known species is badly defined because some specimens attributed to it are in fact white-rayed, glabrate forms of Symphyotrichum puniceum. More work is needed to verify the status of this species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 504. | FNA vol. 20, p. 526. |
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum |
Sibling taxa | ||
Synonyms | Aster anomalus | Aster firmus, Aster lucidulus, Aster puniceus var. firmus |
Name authority | (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) | (Nees) G. L. Nesom: Phytologia 77: 282. (1995) |
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