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manyray aster

Eula's aster, Texas aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, 5–150 cm, colonial; long-rhizomatous (rhizomes thick).
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–3+, erect (straight, stout), glabrous.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

firm, margins serrulate to serrate, distally entire, ± revolute, scabrous, apices mucronate, abaxial faces glabrous or glabrate, adaxial scabrous, cauline often with axillary tufts of leaves;

basal withering by flowering, petiolate (petioles slightly winged, ciliate, sheathing), blades ovate or elliptic to elliptic-lanceolate, 15–120 × 15–35 mm, bases gradually to ± abruptly attenuate, margins crenate-serrate, apices obtuse to acute;

proximal cauline withering by flowering, petiolate to subpetiolate (clasping), blades elliptic to lanceolate, 85–100 × 18–30 mm, bases cuneate to slightly attenuate, apices acute to acuminate;

distal sessile, blades elliptic or elliptic-lanceolate to lanceolate, or lance-oblong to linear-lanceolate (arrays), 8–90 × 1–25 mm, progressively reduced distally, abruptly so in arrays, bases cuneate, margins entire, apices acute to acuminate.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.4–3 cm, secund, glabrous, bracts 5–10, not crowded, linear to linear-lanceolate, spreading, foliaceous, grading into phyllaries.

Involucres

campanulate, 5–10 mm.

hemispheric, (4–)5–6.5 mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

(20–)27–36;

corollas usually white to bluish or lavender-white, rarely pale lavender or lavender-pink, laminae 8–11 × 1–1.5 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

25–31;

corollas yellow turning reddish then brown, (4.1–)4.5–5.8 mm, tubes shorter than funnelform throats, lobes recurved to erect, lanceolate, 1.1–1.3 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 5–6 series, lanceolate to oblong-lanceolate (some inner linear), unequal, bases indurate 1/3–2/3, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate, prominent, apices acute (acuminate), slightly spreading, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in leafy, open, paniculiform arrays, branches ascending to divaricate, nearly horizontal, elongate (to 38 cm), secund-racemiform.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

tan to reddish brown, obovoid to fusiform-obovoid, ± compressed, 1.4–2.9 mm, 4–5-nerved, faces strigillose;

pappi white (sordid), 4.5–5.8 mm.

2n

= 16.

= 48.

Symphyotrichum anomalum

Symphyotrichum eulae

Phenology Flowering late Aug–Oct. Flowering Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams Usually partly shaded, heavy-clayey, clayey-loamy, or silty soils, banks, open stream-bottom woods, ditches, drainage systems, disturbed areas, hedgerows
Elevation 50–500 m (200–1600 ft) 0–100 m (0–300 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum eulae is known only from Texas. It has been reported as a hybrid of S. lanceolatum and S. praealtum. It is not a hybrid, according to Shinners, and we are not considering it as such here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 504. FNA vol. 20, p. 516.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Synonyms Aster anomalus Aster eulae
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Shinners) G. L. Nesom: Phytologia 77: 281. (1995)
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