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manyray aster

Drummond's aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, 30–120 cm, cespitose; short-rhizomatous or with stout, often woody caudices.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–5+, erect (straight, stout), glabrate or glabrescent, sparsely pilose, or hirsuto-pilose proximally, ± densely so distally.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

(abaxially pale or grayish green, adaxially dark green) initially ± firm or membranous, becoming thicker, margins crenate-serrate to serrate, strigoso-ciliate to scabrous, apices acuminate to acute, mucronate, abaxial faces scabrous to densely piloso-strigose, more so on veins, adaxial scabrous to glabrate;

basal withering by flowering, long-petiolate (petioles narrowly winged, sheathing, sometimes ciliate), blades ovate to ovate-oblong or -lanceolate to elliptic or suborbiculate, 10–85(–100) × 8–60 mm, bases deeply cordate to rounded or abruptly attenuate, margins crenate-serrate, apices usually rounded to obtuse, rarely acute;

proximal cauline often withering by flowering, long-petiolate (petioles narrowly winged, more broadly so distally, ciliate), blades ovate or ovate-lanceolate, (25–)50–120(–150) × (15–)20–50(–65) mm, progressively to slightly reduced distally, bases deeply to shallowly cordate or rounded, margins crenate-serrate to sharply serrate;

distal petiolate or subpetiolate or sessile, petioles broadly winged, blades ovate to ovate- or oblong-lanceolate, to oblong-elliptic or linear-lanceolate (4–)5–110 × 1–50 mm, reduced distally (sharply so on array branches in var. texanum), bases shallowly cordate or rounded to attenuate or cuneate (distally), margins crenate-serrate or -serrulate or entire.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0, or 0.2–1.5(–4) cm, densely ascending- or appressed-bracteate, pilosulous, bracts 4–17+, lanceolate or subulate to linear-oblanceolate, distally grading into phyllaries, pilosulous or strigoso-ciliate.

Involucres

campanulate, 5–10 mm.

campanulate, (3.5–)4.5–6(–6.5) mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

(8–)10–15(—20);

corollas usually bright blue to purple or lavender, sometimes white, laminae (5–)7–10(–12) × 1–1.8 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

(10–)13–15(–22);

corollas cream to yellow, becoming deep purple or reddish purple, 3.5–4.5(–5) mm, tubes shorter than funnelform to cylindric throats, lobes triangular to lanceolate, 0.4–0.7 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 4–5 series, subulate or lanceolate (outermost) to narrowly oblong-lanceolate, unequal, bases indurate 1/4–3/4, margins narrowly scarious proximally, hyaline, ciliate, bodies sometimes green to margins distally, green zones lanceolate to linear-lanceolate, apices acute to acuminate or long-acuminate, tips often purple, sometimes involute, mucronulate, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in open, paniculiform arrays, branches ascending to divaricate, often densely racemiform or ± paniculiform (lateral ones open, racemiform) to subsecund, ± stiff to lax, long to relatively short, ± closely to ± remotely leafy, branch leaves progressively or ± abruptly reduced distally.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

dull purple or brown to tan (ribs tan to stramineous), obovoid, ± compressed, (1.5–)2–3 mm, 5–6-nerved, faces glabrate to sparsely strigillose distally;

pappi cream to rose-tinged, 3–4 mm.

2n

= 16.

Symphyotrichum anomalum

Symphyotrichum drummondii

Phenology Flowering late Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams
Elevation 50–500 m (200–1600 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NE; OH; OK; PA; TN; TX; WI; WV; ne Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Variety texanum is tetraploid; var. drummondii is both diploid and tetraploid. The tetraploid race of var. drummondii is most likely autopolyploid; origin of var. texanum has not been determined. The two taxa intergrade and do not appear to warrant species status.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants 40–120 cm; stems proximally sparsely hirsute, distally densely so; leaves firm, basal and proximal long-petiolate, petioles ± winged, proximal bases ± cordate or truncate, sometimes obliquely; heads in ample, paniculiform arrays with divaricate or ascending, bracteate, often racemiform branches; peduncles ± secund, 0–4 cm, bracteate; involucres campanulate, (3.5–)4.5–7 mm; ray corollas usually bright blue, light purple, bluish violet, or lavender, sometimes white; cypselae glabrous
var. drummondii
1. Plants 30–80 cm; stems ± densely hirsute, particularly distally; leaves membranous, becoming thickish, brittle, basal and proximal winged-petiolate, proximal bases cordate, becoming rounded-truncate distally; heads in open, paniculiform arrays with very long, widely spreading branches; peduncles 1–4 cm, densely, minutely bracteate; involucres turbinate to hemispheric, 3.8–5.2 mm; ray corollas bluish white; cypselae strigillose
var. texanum
Source FNA vol. 20, p. 504. FNA vol. 20, p. 503.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. drummondii var. drummondii, S. drummondii var. texanum
Synonyms Aster anomalus Aster drummondii, Aster sagittifolius var. drummondii
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Lindley) G. L. Nesom: Phytologia 77: 279. (1995)
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