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manyray aster

aster ciliolé, fringe blue aster, Lindley's American-aster, Lindley's aster, Lindley's or fringe blue aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, (10–)20–120 cm, colonial or cespitose; usually long-rhizomatous, sometimes with branched caudices.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–3+, erect (straight to ± flexuous), glabrate to sparsely hirsute or strigillose, especially distally.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

thin, margins coarsely, sharply serrate to crenate-serrate or serrulate, ciliate to scabrous, apices acute or acuminate, mucronulate, abaxial faces glabrate to sparsely hirsute, midveins usually densely hirsute, sometimes glabrous, adaxial glabrous or glabrate to scabrellous;

basal usually withering by flowering (sometimes persistent on small plants), long-petiolate (petioles slightly winged, sheathing, ciliate), blades ovate, (24–)40–120(–270) × 15–70 mm, bases usually shallowly cordate, sometimes rounded;

proximal cauline often withering by flowering, winged-petiolate, blades ovate to oblanceolate or lanceolate, (24–)60–150 × (10–)20–60 mm, reduced distally, bases subcordate to cuneate;

distal sessile or sometimes subsessile (petioles widely-winged), blades lance-ovate to linear-lanceolate or linear, 18–75 × (2–)5–25 mm, bases cuneate, sometimes ± clasping, margins serrulate or entire.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.2–1.5+ cm, unequal, reduced distally, ± hirsutulous, bracts 0–4, subulate or linear.

Involucres

campanulate, 5–10 mm.

campanulate, (4–)5–6.5 mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

(12–)14–20;

corollas pale to deep blue or bluish purple, laminae (8.3–)10–15 × 1–2.3 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

14–25(–29);

corollas yellow becoming reddish purple, 4.3–6.4 mm, tubes shorter than funnelform throats, lobes narrowly triangular, 0.5–1.1 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 4–5 series, narrowly oblong-lanceolate to oblong-oblanceolate or linear-oblanceolate (innermost), ± unequal, bases indurate 1/3–2/3, margins narrowly scarious, erose, hyaline or infrequently purplish, sparsely ciliolate, green zones lanceolate to linear-lanceolate or linear, apices acuminate to long-acuminate, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

[(6–)13–50(–100+)] in open, paniculiform arrays, branches ascending.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

yellowish white, obovoid, compressed, 1–2 mm, 5–6-nerved, faces glabrate to sparsely strigillose;

pappi white to pinkish, 3–6 mm.

2n

= 16.

= 48.

Symphyotrichum anomalum

Symphyotrichum ciliolatum

Phenology Flowering late Aug–Oct. Flowering late Jul–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams Rich, open, often ± calcareous, boreal deciduous forests (aspen or aspen-birch-fir-spruce), edges of woods, clearings, aspen or bur oak thickets, sometimes open pine forests, streambanks, trails, roadsides
Elevation 50–500 m (200–1600 ft) 0–2000+ m (0–6600+ ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
IL; MA; ME; MI; MN; MT; ND; NH; NY; SD; VT; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum ciliolatum sporadically hybridizes with S. laeve var. laeve or var. geyeri in their areas of overlap, notably across the Canadian prairies and in south-central Ontario (Manitoulin Island, Bruce Peninsula). It also hybridizes with S. novi-belgii var. novi-belgii in the Gulf of St. Lawrence area [Symphyotrichum ×subgeminatum (Fernald) G. L. Nesom; syn. Aster subgeminatus (Fernald) B. Boivin]. Aster ciliolatus forma comatus Fernald is densely ciliolate along the midnerves and represents an extreme of the variation found in the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 504. FNA vol. 20, p. 502.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Synonyms Aster anomalus Aster ciliolatus, Aster ciliolatus var. comatus, Aster lindleyanus, Aster wilsonii
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Lindley) Á. Löve & D. Löve: Taxon 31: 359. (1982)
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