Symphyotrichum anomalum |
Symphyotrichum ascendens |
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manyray aster |
intermountain aster, long-leaf aster, long-leaf or intermountain or western aster, western American aster, western aster |
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Habit | Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. | Perennials 20–60 cm, colonial; long-rhizomatous. |
Stems | 1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute. |
1–5+, ascending to erect, densely and uniformly strigose, especially distally. |
Leaves | thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils); basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded; proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point; distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate. |
(grayish green) thin to firm, margins entire, apices acute, faces glabrous or strigose; basal usually persistent, petiolate, blades oblanceolate, 50–150 × 5–12(–15) mm, bases attenuate or cuneate, margins entire, apices usually acute; proximal cauline sessile or subpetiolate, blades oblong to narrowly obovate, 50–120 × 5–12 mm, bases attenuate or cuneate; distal sessile, blades 30–70 × 4–10 mm, reduced distally, bases cuneate to ± rounded. |
Peduncles | densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries. |
moderately to densely strigose, bracts 1–4, linear to oblanceolate. |
Involucres | campanulate, 5–10 mm. |
campanulate, 4–7 mm. |
Ray florets | 18–45; corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm. |
(10–)15–40; corollas violet, laminae 8–13 × 1–1.5 mm. |
Disc florets | 33–40+; corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm. |
25–65(–80); corollas yellow, sometimes purple-tinged, 4–6 mm, lobes triangular, 0.4–0.8 mm. |
Phyllaries | in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous. |
in 3–5 series, narrowly oblanceolate or linear (outer) to linear (inner), unequal, bases indurate, margins entire, green zones obovate to elliptic, apices obtuse, faces densely strigose. |
Heads | in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate. |
in paniculiform arrays, branches ascending. |
Cypselae | deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous; pappi tawny or rose-tinged, 3.7–5.2 mm. |
brown, cylindric to obovoid, not compressed, 2–3.5 mm, 3–5-nerved, faces hairy; pappi whitish, 4–7 mm. |
2n | = 16. |
= 26, 36, 52. |
Symphyotrichum anomalum |
Symphyotrichum ascendens |
|
Phenology | Flowering late Aug–Oct. | Flowering Jul–Aug. |
Habitat | Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams | Meadows, grasslands, damp areas in sagebrush steppe and Ponderosa pine woodlands |
Elevation | 50–500 m (200–1600 ft) | 500–3200 m (1600–10500 ft) |
Distribution |
AR; IL; KS; MO; OK
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AZ; CA; CO; ID; MT; ND; NM; NV; OR; UT; WA; WY; AB; BC; SK
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Discussion | A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum ascendens is widely distributed in the Great Basin. It is an allopolyploid derived from the hybrid between S. spathulatum (x = 8) and S. falcatum (x = 5). Chromosome numbers differ markedly in their geographic distribution, 2n = 26 prevailing in the southwestern part of the range, and 2n = 52 in the northeastern part (G. A. Allen 1985) Backcrosses to both parental species or hybrids with related taxa are sometimes seen where the ranges overlap. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 504. | FNA vol. 20, p. 538. |
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | Asteraceae > tribe Astereae > Symphyotrichum > subg. Ascendentes |
Sibling taxa | ||
Synonyms | Aster anomalus | Aster ascendens, Aster chilensis subsp. ascendens |
Name authority | (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) | (Lindley) G. L. Nesom: Phytologia 77: 276. (1995) |
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