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Sullivant's coolwort

coolwort, sullivantia

Habit Plants not stoloniferous. Herbs, rhizomatous, stoloniferous or not; caudex not cormlike, with persistent leaf bases.
Flowering stems

5–40 cm, erect to ascending.

erect to ascending, leafy or leafless, 5–60 cm, glabrate or sparsely stipitate-glandular.

Leaves

blade 1–12 cm wide, 1–2 times toothed, 7–13-lobed for 1/4–1/2 length to shallowly and somewhat irregularly 7–25-lobed;

lobes cuneate-oblong to crenate and 1–2 times denticulate to coarsely subcrenulate, margins erose with long bristles to lacinate.

in basal rosette and cauline;

cauline leaves similar to basal leaves, becoming smaller and short-petiolate to sessile distally;

stipules present;

petiole glabrate or sparsely stipitate-glandular;

blade reniform to reniform-orbiculate, 5–25-lobed, base cordate in basal leaves, cordate to truncate in cauline leaves, margins 1–2 times dentate to crenate, apex rounded, surfaces glabrate;

venation palmate.

Inflorescences

erect to decumbent, primary and higher-order branches ascending relative to central axis.

diffuse thyrses of monochasial or dichasial, compound cymes, terminal from terminal bud in rosette, 10–70-flowered, bracteate, (stipitate-glandular, especially distally, slightly viscid with sweet, resinous odor).

Flowers

hypanthium tubular-campanulate, portion distal to adnation to ovary not marked by conspicuous flaring, 1.3–3.2 × 1.2–2.1 mm, stipitate-glandular proximally;

sepals triangular-ovate to lanceolate-oblong, not as broad as long at apex, 0.8–1.5 × 0.6–1.1 mm, apex acute;

petals unlobed, 1.7–3.5 mm, ovate and abruptly narrowed in claw to lanceolate and gradually narrowed into claw, 0.7–1.6 mm wide;

ovary ca. 1/2 inferior.

(lax and pendent at anthesis), (pedicellate);

hypanthium adnate to ovary 1/3–1/2 proximal length, free from ovary to 1.5 mm, green;

sepals 5, green;

petals 5, white;

nectary disc absent;

stamens 5, (shorter than sepals);

filaments subulate, (± as long as anthers); (anthers yellow);

ovary 1/2–4/5+ inferior at anthesis, 2-locular, carpels connate proximally;

placentation axile;

styles 2, (erect);

stigmas 2.

Capsules

cylindric-ovate to cylindric-elliptic, 3–7.5 × 1.3–3.2 mm, beaks ± spreading.

2-beaked.

Seeds

usually tan, rarely red-brown, 0.7–1.2 × 0.2–0.5 mm, main body (excluding winged margins) 0.4–0.7 mm.

(10–100), tan or dark red-brown, linear-fusiform, (narrowly wing-margined), smooth.

x

= 7.

2n

= 14.

Sullivantia sullivantii

Sullivantia

Phenology Flowering May–Jul.
Habitat Moist limestone and sandstone cliffs
Elevation 300-700 m (1000-2300 ft)
Distribution
from FNA
IA; IL; IN; KY; MN; MO; OH; TN; VA; WI
[WildflowerSearch map]
[BONAP county map]
from USDA
United States
[BONAP county map]
Discussion

Species 3 (3 in the flora).

Perhaps the most noteworthy features of the distribution of Sullivantia species are their disjunct distributions and the restriction of most populations to unglaciated areas near Pleistocene glacial margins (D. E. Soltis 1991).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Hypanthia tubular-campanulate, portion distal to adnation to ovary not marked by conspicuous flaring; primary and higher-order inflorescence branches ascending relative to central axis; ovaries ca. 1/2 inferior; seeds usually tan, rarely red-brown, main body (excluding winged margins) 0.4-0.7 mm.
S. sullivantii
1. Hypanthia turbinate-campanulate, portion distal to adnation to ovary marked by conspicuous flaring; primary and higher-order inflorescence branches usually perpendicular to central axis; ovaries 4/5+ inferior; seeds dark red-brown, main body (excluding winged margins) 0.7-1 mm
→ 2
2. Plants stoloniferous; petal blades suborbiculate; hypanthia widely turbinate-campanulate, sepals broader than long at apex.
S. oregana
2. Plants not stoloniferous; petal blades ovate to ovate-lanceolate; hypanthia narrowly turbinate-campanulate, sepals not as broad as long at apex.
S. hapemanii
Source FNA vol. 8, p. 122. FNA vol. 8, p. 121. Author: Douglas E. Soltis.
Parent taxa Saxifragaceae > Sullivantia Saxifragaceae
Sibling taxa
S. hapemanii, S. oregana
Subordinate taxa
S. hapemanii, S. oregana, S. sullivantii
Synonyms Saxifraga sullivantii, Saxifraga renifolia
Name authority (Torrey & A. Gray) Britton: Mem. Torrey Bot. Club 5: 178. (1894) Torrey & A. Gray: Amer. J. Sci. Arts 42: 22. 1842 ,
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