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coolwort, sullivantia

Habit Herbs, rhizomatous, stoloniferous or not; caudex not cormlike, with persistent leaf bases.
Flowering stems

erect to ascending, leafy or leafless, 5–60 cm, glabrate or sparsely stipitate-glandular.

Leaves

in basal rosette and cauline;

cauline leaves similar to basal leaves, becoming smaller and short-petiolate to sessile distally;

stipules present;

petiole glabrate or sparsely stipitate-glandular;

blade reniform to reniform-orbiculate, 5–25-lobed, base cordate in basal leaves, cordate to truncate in cauline leaves, margins 1–2 times dentate to crenate, apex rounded, surfaces glabrate;

venation palmate.

Inflorescences

diffuse thyrses of monochasial or dichasial, compound cymes, terminal from terminal bud in rosette, 10–70-flowered, bracteate, (stipitate-glandular, especially distally, slightly viscid with sweet, resinous odor).

Flowers

(lax and pendent at anthesis), (pedicellate);

hypanthium adnate to ovary 1/3–1/2 proximal length, free from ovary to 1.5 mm, green;

sepals 5, green;

petals 5, white;

nectary disc absent;

stamens 5, (shorter than sepals);

filaments subulate, (± as long as anthers); (anthers yellow);

ovary 1/2–4/5+ inferior at anthesis, 2-locular, carpels connate proximally;

placentation axile;

styles 2, (erect);

stigmas 2.

Capsules

2-beaked.

Seeds

(10–100), tan or dark red-brown, linear-fusiform, (narrowly wing-margined), smooth.

x

= 7.

Sullivantia

Distribution
from USDA
United States
[BONAP county map]
Discussion

Species 3 (3 in the flora).

Perhaps the most noteworthy features of the distribution of Sullivantia species are their disjunct distributions and the restriction of most populations to unglaciated areas near Pleistocene glacial margins (D. E. Soltis 1991).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Hypanthia tubular-campanulate, portion distal to adnation to ovary not marked by conspicuous flaring; primary and higher-order inflorescence branches ascending relative to central axis; ovaries ca. 1/2 inferior; seeds usually tan, rarely red-brown, main body (excluding winged margins) 0.4-0.7 mm.
S. sullivantii
1. Hypanthia turbinate-campanulate, portion distal to adnation to ovary marked by conspicuous flaring; primary and higher-order inflorescence branches usually perpendicular to central axis; ovaries 4/5+ inferior; seeds dark red-brown, main body (excluding winged margins) 0.7-1 mm
→ 2
2. Plants stoloniferous; petal blades suborbiculate; hypanthia widely turbinate-campanulate, sepals broader than long at apex.
S. oregana
2. Plants not stoloniferous; petal blades ovate to ovate-lanceolate; hypanthia narrowly turbinate-campanulate, sepals not as broad as long at apex.
S. hapemanii
Source FNA vol. 8, p. 121. Author: Douglas E. Soltis.
Parent taxa Saxifragaceae
Subordinate taxa
S. hapemanii, S. oregana, S. sullivantii
Name authority Torrey & A. Gray: Amer. J. Sci. Arts 42: 22. 1842 ,
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