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mesquite neststraw, Sonoran neststraw

neststraw, stylocline

Habit Plants 2–10(–15) cm. Annuals, 1–10(–20) cm.
Stems

1, ± erect, or 2–10+, ascending to ± prostrate.

Leaves

blunt (proximal) or acute (median and distal), mucronate, longest 6–13 mm;

largest capitular leaves ± elliptic to narrowly ovate, 3–10 × 2–3 mm.

cauline; mostly alternate;

blades oblanceolate to lanceolate or broader.

Involucres

0 or inconspicuous.

Receptacles

clavate, 1.2–2.2 mm, heights 2.8–3.5 times diams.;

scars ± evenly distributed, mamillate.

cylindric to clavate (heights 2.8–8 times diams.), glabrous.

Pistillate florets

12–25+.

Bisexual florets

0.

Phyllaries

0, vestigial, or falling, ± subulate, mostly 0.1–0.5 mm, unequal.

0, vestigial, or 1–4, unequal or subequal (similar to paleae).

Heads

in cymiform to ± paniculiform, sometimes dichasiform arrays, ± spheric, largest 3.5–4.5 × 3–4 mm, thickly lanuginose.

in glomerules of 2–10 (rarely borne singly) in ± dichasiform (sometimes ± paniculiform) arrays.

Cypselae

0.6–0.8 mm, slightly compressed;

pappi: staminate of (1–)3–8 barbellate bristles 0.9–1.3 mm (proximal barbs longer, spreading).

brown, monomorphic: compressed to obcompressed, ± obovoid, slightly incurved, abaxially gibbous, faces glabrous, smooth, shiny;

corolla scars subapical;

pappi: pistillate 0, staminate rarely 0, usually of 1–10(–13) bristles (hidden in heads).

Pistillate

paleae: longest 1.9–3.1 mm, winged distally;

wings ± elliptic, widest in distal 1/3 of palea lengths;

bodies (except midnerves) chartaceous;

outermost paleae ± saccate.

paleae falling, erect to ascending;

bodies with 5+ nerves (nerves ± parallel, obscure), ± ovate or boat-shaped, saccate most of lengths (obcompressed to terete, not galeate, each usually enclosing a floret, outermost open in 2 spp.);

wings ± erect (apical).

Functionally

staminate florets 2–5;

ovaries partially developed, 0.3–0.6 mm;

corollas 0.9–1.4 mm.

staminate florets 2–6;

corolla lobes (4–)5, ± equal.

Staminate

paleae readily falling, mostly 2–4, erect in fruit (not enlarged), shorter than pistillate paleae;

bodies linear-lanceolate to oblanceolate.

x

= 14.

Stylocline sonorensis

Stylocline

Phenology Flowering and fruiting Mar–May.
Habitat Grassy hillsides, sandy drainages, with mesquite (Prosopis)
Elevation 400–1400 m (1300–4600 ft)
Distribution
from FNA
AZ; CA; Mexico (Sonora)
[BONAP county map]
from USDA
sw United States; nw Mexico
[BONAP county map]
Discussion

Stylocline sonorensis is known from southeastern Arizona and northeastern Sonora. A disjunct, 1930 California occurrence was from apparently suitable habitat; recent searches have not relocated it.

Stylocline sonorensis is illustrated in J. D. Morefield (1992). It is superficially similar to S. citroleum; its closest relative is S. micropoides. Its ancestors may have been hybrid products involving Logfia depressa or its progenitors (Morefield).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 7 (7 in the flora).

See discussion of Filagininae following the tribal description (p. 385).

Stylocline occurs in Mediterranean, desert, and semi-desert climates; most species appear only after exceptionally wet winters, or in moisture-accumulating microsites (e.g., rock bases, washes, shrub drip-lines). Plants usually grow in undisturbed soils (often with soil crusts) and sometimes colonize stabilized disturbances.

In some species of Stylocline, the outermost bracts of heads are merely concave, not saccate; these are paleae (if they subtend and fall with florets) or phyllaries (if they persist and subtend only adjacent saccate paleae). Texture of the palea bodies is diagnostic for each species. In dried specimens, chartaceous bodies tear easily and irregularly when the abaxial indument is gently scraped. Cartilaginous bodies can be scraped clean without tearing and split lengthwise only if forced.

Stylocline appears to be ancestral to Micropus and Psilocarphus, and derived from, sister to, and/or reticulate with Logfia (J. D. Morefield 1992). Stylocline citroleum, S. sonorensis, and L. depressa show some transitional traits between the genera.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Longest pistillate paleae winged proximally and distally, wings widest in proximal 2/3 of palea lengths; phyllaries ± persistent, elliptic or broader, 1–3.5 mm
→ 2
1. Longest pistillate paleae winged distally, wings widest in distal 1/3 of palea lengths; phyllaries 0, vestigial, or falling, ± subulate, mostly 0.1–0.5 mm
→ 3
2. Longest pistillate paleae: wings broadly ovate (bases rounded or cordate); staminate ovaries vestigial, 0–0.2 mm, pappi usually of 1–5 bristles; heads arachnoid to thinly lanuginose (often shiny, indument obscured by palea wings)
S. gnaphaloides
2. Longest pistillate paleae: wings elliptic to slightly obovate (bases acute); staminate ovaries partially developed, (0.2–)0.3–0.6 mm, pappi of (5–)6–12(–13) bristles; heads thickly lanuginose (dull, indument evident)
S. citroleum
3. Receptacles clavate, heights 2.8–3.5 times diams.; staminate ovaries partially developed, 0.3–0.6 mm (cypselae 0.6–0.8 mm; heads ± spheric, diams. 3–4 mm; longest pistillate paleae 1.9–3.1 mm; proximal leaves blunt)
S. sonorensis
3. Receptacles ± cylindric, heights 4–8 times diams.; staminate ovaries ± vestigial, 0–0.3(–0.4) mm (cypselae 0.8–1.6 mm, heads ovoid to ellipsoid or diams. 5–9 mm, longest pistillate paleae 3.4–4.5 mm and/or proximal leaves acute)
→ 4
4. Heads ± spheric, thickly lanuginose, largest diams. 5–9 mm; pistillate paleae: longest 3.4–4.5 mm, outermost saccate
→ 5
4. Heads ovoid to ellipsoid, thinly lanuginose, largest diams. 1.5–4 mm; pistillate paleae: longest 2–3.3 mm, outermost open, concave
→ 6
5. Bodies of longest pistillate paleae (except midnerves) chartaceous; cypselae compressed; largest capitular leaves (some or all) subulate to lanceolate (widest in proximal 1/3), (7–)11–17 mm (distalmost mainly 1.5–2 times head heights)
S. micropoides
5. Bodies of longest pistillate paleae cartilaginous; cypselae obcompressed; largest capitular leaves (all) ± elliptic to ± oblanceolate (widest in distal 2/3), 4–11 mm (distalmost mainly 0.8–1.2 times head heights)
S. intertexta
6. Heads 2.5–4 mm diam.; longest pistillate paleae 2.8–3.3 mm; cypselae 1.1–1.6 mm; staminate corollas 1.1–1.7 mm (lobes usually 5); leaves ± acute
S. psilocarphoides
6. Heads 1.5–2.5 mm diam.; longest pistillate paleae 2–2.7 mm; cypselae 0.7–1 mm; staminate corollas 0.8–1.1 mm (lobes usually 4); leaves blunt
S. masonii
Source FNA vol. 19, p. 452. FNA vol. 19, p. 450. Author: James D. Morefield.
Parent taxa Asteraceae > tribe Gnaphalieae > Stylocline Asteraceae > tribe Gnaphalieae
Sibling taxa
S. citroleum, S. gnaphaloides, S. intertexta, S. masonii, S. micropoides, S. psilocarphoides
Subordinate taxa
S. citroleum, S. gnaphaloides, S. intertexta, S. masonii, S. micropoides, S. psilocarphoides, S. sonorensis
Name authority Wiggins: Contr. Dudley Herb. 4: 26. (1950) Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 338. (1840)
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