Strophostyles |
Strophostyles umbellata |
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fuzzy bean |
perennial wildbean, perennial woolly bean, pink fuzzy or perennial woolly or perennial sand bean, pink fuzzybean |
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Stems | terete, often lignescent at base, from subterranean, branched caudex, sometimes with adventitious roots. |
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Leaves | alternate, odd-pinnate, pulvinate; stipules present, persistent, sessile, divergent from stem, striate, triangular-ovate, not produced proximal to point of insertion; long-petiolate, petiole and rachis canaliculate, pubescent; leaflets 3, stipels persistent, striate, often curved, linear, blades 1.3–5.6(–7.2) cm, membranous to papery, margins entire or lobed, sometimes basally lobed, surfaces sericeous or strigose. |
leaflet blades usually ovate-lanceolate, sometimes narrowly lanceolate, margins entire or shallowly, rarely deeply, lobed, sinus 0–1(–6) mm, lateral leaflets 1.5–4.2 × 0.3–2.7 cm, terminal leaflet (1.5–)2–4(–7) × 0.2–2.2(–3) cm, surfaces strigose. |
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Inflorescences | 1–12(–22)-flowered, axillary, pseudoracemes, rachis much contracted, with 1–6(–11)-flowering nodes, nodes often swollen, glandular, each bearing at most 2 flowers; primary bracts early deciduous or absent; secondary bracts subtending pedicels, persistent, ovate to lanceolate; bracteoles persistent, rigid and striate, lanceolate, equal to or longer than calyx tube. |
with (1 or)2–6(–11) flower-bearing nodes. |
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Peduncles | angular. |
stout, lignescent, (4.4–)6–30 cm, 0.8–1.3 mm diam. |
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Pedicels | mostly shorter than calyx tube, thickened in fruit. |
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Flowers | papilionaceous; calyx campanulate, 1.3–7 mm, lobes 4 (due to connation of 2 adaxial lobes into 1 acute-attenuate lobe), abaxial lobe more narrowly triangular and often longest, lateral lobes triangular, adaxial lobes connate into acute lobe; corolla pink or pinkish, banner often with yellowish maculae, keel pinkish, tip (beak) dark purple, 3.6–15 mm; wings oblong, not projected beyond distal bend of keel; keel petals connate along upper margin, incurved, with a prominent gibbosity along upper margin proximal to beak; stamens 10, diadelphous; anthers sub-basifixed, uniform; pollen tricolporate; ovary sessile, arched, sparsely to densely strigose, at base surrounded by a nectariferous sheath; style incurved like the keel, often becoming twisted, jointed at the first (proximal) curve, distinctly incrassate and flattened, distal portion introrsely bearded; stigma terminal and introrse. |
bracteoles 0.8–2.4 mm, shorter than calyx tube; calyx tube (1.4–)2–4 mm, lobes 1–2.5(–3) mm; banner pink, 7.3–15 mm; wings pinkish, 7–12 mm; keel 8–13 mm, beak prominent, erect, slightly curved, 1.5–2 mm diam., closely positioned to face of banner petal, protruding well above wings. |
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Fruits | legumes, sessile, held horizontally or somewhat drooping, linear or slightly curved, cylindric to subcylindric, dehiscent with twisting valves, sericeous, strigose, glabrate, or glabrous. |
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Legumes | subcylindric, laterally compressed, 3–7 × 0.2–0.6 cm, glabrate or sparsely strigose. |
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Seeds | 4–10, spherical or oblong, subquadrate or reniform, ends truncate, usually covered with a cellular layer; hilum oblanceolate, elongated 1/2 length of seed, rim-aril and hilar tongue much reduced, lens distinct and divided; seedlings with epigeal germination. |
5–10, sometimes faintly mottled, with cellular or waxy covering. |
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Vines | , annual or perennial, unarmed, to 2 m. Stems slender, flexible, trailing or climbing, often branching at proximal nodes, not woody, sometimes lignescent at base, sparsely to densely short-pilose. |
perennial, with slender taproot. |
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x | = 11. |
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2n | = 22. |
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Strophostyles |
Strophostyles umbellata |
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Phenology | Flowering Jul–Sep. | |||||||||
Habitat | Fields, meadows, rarely in woodlands. | |||||||||
Elevation | 0–1500 m. (0–4900 ft.) | |||||||||
Distribution |
North America; n Mexico |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NJ; NM; NY; OK; PA; RI; SC; SD; TN; TX; VA; WI; WV
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Discussion | Species 3 (3 in the flora). Strophostyles flowers fade to pale brownish yellow when dry; the keel is proximally broad, with a prominent gibbosity along the upper margin proximal to the beak, usually with a longitudinal fold where it is adherent to the wing, gradually narrowing above this; the tubular apical portion curves to one side of the flower. Systematics and phylogenetic relationships of Strophostyles have been studied (E. T. Riley-Hulting et al. 2004). Additional clarification on the nomenclature of Strophostyles have recently been published (A. Delgado-Salinas and M. Lavin 2004). Strophostyles species have been reported to serve as food and to have medicinal properties (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Strophostyles umbellata is highly variable in leaflet shape, from narrowly lanceolate like those of S. leiosperma to ovate-lanceolate like those of S. helvola. It is best distinguished by its prominent keel beak, which has a thick base, a curved tip, and is held close to the banner face. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 11. | FNA vol. 11. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Phasellus, Phaseolus section strophostyles | Glycine umbellata, Phaseolus umbellatus, S. umbellata var. paludigena | ||||||||
Name authority | Elliott: Sketch Bot. S. Carolina 2: 229. (1823) — name conserved | (Muhlenberg ex Willdenow) Britton in N. L. Britton and A. Brown: Ill. Fl. N. U.S. 2: 339. (1897) | ||||||||
Web links |