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fuzzy bean

perennial wildbean, perennial woolly bean, pink fuzzy or perennial woolly or perennial sand bean, pink fuzzybean

Stems

terete, often lignescent at base, from subterranean, branched caudex, sometimes with adventitious roots.

Leaves

alternate, odd-pinnate, pulvinate;

stipules present, persistent, sessile, divergent from stem, striate, triangular-ovate, not produced proximal to point of insertion; long-petiolate, petiole and rachis canaliculate, pubescent;

leaflets 3, stipels persistent, striate, often curved, linear, blades 1.3–5.6(–7.2) cm, membranous to papery, margins entire or lobed, sometimes basally lobed, surfaces sericeous or strigose.

leaflet blades usually ovate-lanceolate, sometimes narrowly lanceolate, margins entire or shallowly, rarely deeply, lobed, sinus 0–1(–6) mm, lateral leaflets 1.5–4.2 × 0.3–2.7 cm, terminal leaflet (1.5–)2–4(–7) × 0.2–2.2(–3) cm, surfaces strigose.

Inflorescences

1–12(–22)-flowered, axillary, pseudoracemes, rachis much contracted, with 1–6(–11)-flowering nodes, nodes often swollen, glandular, each bearing at most 2 flowers;

primary bracts early deciduous or absent;

secondary bracts subtending pedicels, persistent, ovate to lanceolate;

bracteoles persistent, rigid and striate, lanceolate, equal to or longer than calyx tube.

with (1 or)2–6(–11) flower-bearing nodes.

Peduncles

angular.

stout, lignescent, (4.4–)6–30 cm, 0.8–1.3 mm diam.

Pedicels

mostly shorter than calyx tube, thickened in fruit.

Flowers

papilionaceous;

calyx campanulate, 1.3–7 mm, lobes 4 (due to connation of 2 adaxial lobes into 1 acute-attenuate lobe), abaxial lobe more narrowly triangular and often longest, lateral lobes triangular, adaxial lobes connate into acute lobe;

corolla pink or pinkish, banner often with yellowish maculae, keel pinkish, tip (beak) dark purple, 3.6–15 mm;

wings oblong, not projected beyond distal bend of keel;

keel petals connate along upper margin, incurved, with a prominent gibbosity along upper margin proximal to beak;

stamens 10, diadelphous;

anthers sub-basifixed, uniform;

pollen tricolporate;

ovary sessile, arched, sparsely to densely strigose, at base surrounded by a nectariferous sheath;

style incurved like the keel, often becoming twisted, jointed at the first (proximal) curve, distinctly incrassate and flattened, distal portion introrsely bearded;

stigma terminal and introrse.

bracteoles 0.8–2.4 mm, shorter than calyx tube;

calyx tube (1.4–)2–4 mm, lobes 1–2.5(–3) mm;

banner pink, 7.3–15 mm;

wings pinkish, 7–12 mm;

keel 8–13 mm, beak prominent, erect, slightly curved, 1.5–2 mm diam., closely positioned to face of banner petal, protruding well above wings.

Fruits

legumes, sessile, held horizontally or somewhat drooping, linear or slightly curved, cylindric to subcylindric, dehiscent with twisting valves, sericeous, strigose, glabrate, or glabrous.

Legumes

subcylindric, laterally compressed, 3–7 × 0.2–0.6 cm, glabrate or sparsely strigose.

Seeds

4–10, spherical or oblong, subquadrate or reniform, ends truncate, usually covered with a cellular layer;

hilum oblanceolate, elongated 1/2 length of seed, rim-aril and hilar tongue much reduced, lens distinct and divided;

seedlings with epigeal germination.

5–10, sometimes faintly mottled, with cellular or waxy covering.

Vines

, annual or perennial, unarmed, to 2 m. Stems slender, flexible, trailing or climbing, often branching at proximal nodes, not woody, sometimes lignescent at base, sparsely to densely short-pilose.

perennial, with slender taproot.

x

= 11.

2n

= 22.

Strophostyles

Strophostyles umbellata

Phenology Flowering Jul–Sep.
Habitat Fields, meadows, rarely in wood­lands.
Elevation 0–1500 m. (0–4900 ft.)
Distribution
from USDA
North America; n Mexico
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NJ; NM; NY; OK; PA; RI; SC; SD; TN; TX; VA; WI; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 3 (3 in the flora).

Strophostyles flowers fade to pale brownish yellow when dry; the keel is proximally broad, with a prominent gibbosity along the upper margin proximal to the beak, usually with a longitudinal fold where it is adherent to the wing, gradually narrowing above this; the tubular apical portion curves to one side of the flower.

Systematics and phylogenetic relationships of Strophostyles have been studied (E. T. Riley-Hulting et al. 2004). Additional clarification on the nomenclature of Strophostyles have recently been published (A. Delgado-Salinas and M. Lavin 2004). Strophostyles species have been reported to serve as food and to have medicinal properties (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Strophostyles umbellata is highly variable in leaflet shape, from narrowly lanceolate like those of S. leiosperma to ovate-lanceolate like those of S. helvola. It is best distinguished by its prominent keel beak, which has a thick base, a curved tip, and is held close to the banner face.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Keel petals: beak mostly concealed by wings; peduncles slender, herbaceous, 1.2–11(–12.3) cm in fruit; leaf blades and fruits sericeous; banners 3.6–7(–8.3) mm; legumes 1.2–4.7 cm; leaflet blades lanceolate, margins usually entire, never deeply lobed.
S. leiosperma
1. Keel petals: beak protruding well above wings; peduncles stout, lignescent, (2.4–)5–30 cm in fruit; leaf blades and fruits sparsely strigose to glabrate or glabrous; banners 6.7–15 mm; legumes 3–7(–9.6) cm; leaflet blades lanceolate to narrowly lanceolate or pandurate, margins entire or lobed.
→ 2
2. Keel petals with slender, curved beaks; bracteoles 1.5–4.2 mm, equal to or longer than calyx tube; legumes cylindric; leaflet blade margins lobed or entire, terminal leaflet blades 1.8–7.2 × 0.9–4.6 cm.
S. helvola
2. Keel petals with stout, erect, slightly curved beaks; bracteoles 0.8–2.4 mm, shorter than calyx tube; legumes subcylindric; leaflet blade margins usually entire or shallowly lobed, rarely deeply lobed, terminal leaflet blades (1.5–)2–4(–7) × 0.2–2.2(–3) cm.
S. umbellata
Source FNA vol. 11. Authors: Alfonso Delgado-Salinas, Matt Lavin, Erin Thais Riley. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae Fabaceae > subfam. Faboideae > Strophostyles
Sibling taxa
S. helvola, S. leiosperma
Subordinate taxa
S. helvola, S. leiosperma, S. umbellata
Synonyms Phasellus, Phaseolus section strophostyles Glycine umbellata, Phaseolus umbellatus, S. umbellata var. paludigena
Name authority Elliott: Sketch Bot. S. Carolina 2: 229. (1823) — name conserved (Muhlenberg ex Willdenow) Britton in N. L. Britton and A. Brown: Ill. Fl. N. U.S. 2: 339. (1897)
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