The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

milkwort jewelflower

jewelweed, mountain jewel-flower, shield plant, shieldleaf

Habit Annuals; (sometimes glaucous), glabrous throughout. Biennials or perennials; (short-lived); (glaucous), glabrous throughout.
Stems

unbranched or branched distally, (0.8–)2–8(–10) dm.

unbranched or branched (several) basally and distally, (0.5–)1.5–12(–15) dm.

Basal leaves

(soon withered); rosulate;

petiolate;

blade 1- or 2-pinnatifid (with broadly linear to filiform lobes), 2–20 cm, margins sinuate-dentate.

(soon withered);

subrosulate;

petiolate;

blade broadly ovate, obovate or oblong, 1.5–6 cm, margins entire or repand, or dentate apically.

Cauline leaves

blade linear, 1–10 cm × 1–3 mm, (smaller distally), base auriculate, margins entire.

blade oblong to obovate or suborbicular, or (distally) orbicular or oblong-ovate, (0.7–)1.5–6(–9) cm × (4–)10–45 mm, base auriculate to amplexicaul (amplexicaul distally), margins entire or repand.

Racemes

ebracteate, (lax).

bracteate below or between proximalmost 1 or 2 flowers, (lax to compact, secund or not).

Flowers

(markedly zygomorphic);

calyx urceolate;

sepals greenish yellow or purplish, (abaxial broadly ovate, not keeled, 4–6 × 3–4 mm, lateral ovate-lanceolate, keeled, 4–6 × 1.5–2 mm, adaxial suborbicular to broadly ovate-cordate, forming a bannerlike hood, keeled, 4–6 × 6–8 mm);

petals white (with brownish veins), 5–8 mm, blade 1–3 × 0.7–1.2 mm, margins crisped (channeled), claw 4–5 mm, wider than blade;

stamens in 3 unequal pairs;

filaments: abaxial pair (distinct), 4–5 mm, lateral pair 3–4 mm, adaxial pair (connate), 5–6 mm;

anthers: abaxial and lateral pairs fertile, 1.5–2 mm, adaxial pairs sterile, 0.3–0.7 mm;

gynophore 0.1–0.4 mm.

calyx urceolate;

sepals purplish, gray-green, or yellowish, 6–10(–13) mm, keeled or not, (apex recurved);

petals purplish or yellowish white (usually with purple veins), (6–)8–14 mm, blade 2–5 × 1–2.5 mm, margins not crisped, claw 4–10 mm, as wide as or wider than blade, (apex reflexed);

stamens in 3 unequal pairs;

filaments (distinct): abaxial pair (2.5–)4–7 mm, lateral pair (1.5–)3–5 mm, adaxial pair (5–)7–11 mm;

anthers (all) fertile or adaxial pair with reduced fertility and shorter, (1.5–)2.5–4.5(–6) mm;

gynophore 0.2–1 mm.

Fruiting pedicels

strongly recurved, 2–5 mm.

divaricate to ascending, (2–)3–12(–17) mm, (expanded at receptacle).

Fruits

pendent, smooth, straight, flattened, 2.4–5.6 cm × 1.2–1.7 mm;

valves each with obscure or somewhat prominent midvein;

replum straight;

ovules (10–)18–50 per ovary;

style 0.8–2 mm;

stigma entire.

arcuate-spreading to pendent, torulose or smooth, straight, flattened, (3–)4–13(–16) cm × 1.5–2.5(–3) mm;

valves each with obscure or somewhat prominent midvein;

replum straight;

ovules 26–76(–110) per ovary;

style 0.4–0.7 mm;

stigma entire.

Seeds

oblong, 1.7–2 × 0.9–1.1 mm;

wing 0.2–0.3 mm wide, distal.

broadly oblong to ovoid or orbicular, 1.5–2.5 × 0.8–1.7 mm;

wing 0.1–0.5 mm wide, continuous or on distal 1/2.

2n

= 28.

= 28.

Streptanthus polygaloides

Streptanthus tortuosus

Phenology Flowering May–Jul. Flowering Apr–Sep.
Habitat Serpentine substrates in grasslands, openings chaparral, oak and pine woodlands Rocky open slopes, open woodlands, montane forests, alpine areas
Elevation 200-1900 m (700-6200 ft) 200-4100 m (700-13500 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Streptanthus polygaloides is highly variable in plant height, flower color, fruit size, and dissection of basal leaves. Further studies may lead to recognition of infraspecific taxa. The species is distributed in Butte, Calaveras, El Dorado, Fresno, Mariposa, Nevada, Sierra, Tuolumne, and Yuba counties.

Streptanthus polygaloides is one of the few nickel hyperaccumulators in the flora area, and it averages 2,430–18,600 µg/g dry weight (R. D. Reeves et al. 1981; A. R. Kruckeberg and Reeves 1995).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Streptanthus tortuosus is highly polymorphic, forming distinct local races. Variation in duration, stature, branching, development of the sepal keel, petal color, and bract size are not correlated with habitat or geography. Some authors (e.g., R. C. Rollins 1993; R. E. Buck et al. 1993) divided it into five varieties based on variations in those characters, but the variation is continuous and the purported differences do not hold. For example, those authors and N. H. Holmgren (2005b), recognized the highly branched, shorter plants of the alpine Sierra Nevada as var. orbiculatus, but intergradation downslope is completely clinal. The most distinctive variant (S. foliosus) is restricted to the central Sierra Nevada, where plants with large, brittle, subachlorophyllus bracts predominate. However, that condition reappears sporadically in the Klamath Ranges and elsewhere. Annual plants have never been documented in S. tortuosus, contrary to the claim by some authors (e.g., Rollins; Buck et al.; Holmgren); the alleged “annuals” produce rosettes that invariably overwinter before flowering. Without thorough molecular, experimental, and cytological study of this complex, it is impractical to recognize ill-defined varieties that represent only a minor fraction of the overall variation in the species.

R. C. Rollins (1993) placed Streptanthus foliosus in the synonymy of S. diversifolius, but its type clearly belongs to S. tortuosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 720. FNA vol. 7, p. 722.
Parent taxa Brassicaceae > tribe Thelypodieae > Streptanthus Brassicaceae > tribe Thelypodieae > Streptanthus
Sibling taxa
S. barbatus, S. barbiger, S. batrachopus, S. bernardinus, S. brachiatus, S. bracteatus, S. breweri, S. callistus, S. campestris, S. carinatus, S. cordatus, S. cutleri, S. diversifolius, S. drepanoides, S. farnsworthianus, S. fenestratus, S. glandulosus, S. gracilis, S. hesperidis, S. hispidus, S. howellii, S. hyacinthoides, S. insignis, S. longisiliquus, S. maculatus, S. morrisonii, S. oblanceolatus, S. oliganthus, S. petiolaris, S. platycarpus, S. squamiformis, S. tortuosus, S. vernalis, S. vimineus
S. barbatus, S. barbiger, S. batrachopus, S. bernardinus, S. brachiatus, S. bracteatus, S. breweri, S. callistus, S. campestris, S. carinatus, S. cordatus, S. cutleri, S. diversifolius, S. drepanoides, S. farnsworthianus, S. fenestratus, S. glandulosus, S. gracilis, S. hesperidis, S. hispidus, S. howellii, S. hyacinthoides, S. insignis, S. longisiliquus, S. maculatus, S. morrisonii, S. oblanceolatus, S. oliganthus, S. petiolaris, S. platycarpus, S. polygaloides, S. squamiformis, S. vernalis, S. vimineus
Synonyms Microsemia polygaloides Disaccanthus tortuosus, Erysimum tortuosum, Pleiocardia foliosa, Pleiocardia orbiculata, Pleiocardia suffrutescens, Pleiocardia tortuosa, S. foliosus, S. orbiculatus, S. sanhedrensis, S. suffrutescens, S. tortuosus var. flavescens, S. tortuosus var. oblongus, S. tortuosus var. optatus, S. tortuosus var. orbiculatus, S. tortuosus var. pallidus, S. tortuosus var. suffrutescens
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 519. (1865) Kellogg: Proc. Calif. Acad. Sci. 2: 152, plate 46. (1863)
Web links