Streptanthus polygaloides |
Streptanthus insignis |
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milkwort jewelflower |
plumed jewelflower, San Benito jewelflower |
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Habit | Annuals; (sometimes glaucous), glabrous throughout. | Annuals; hirsute-hispid throughout. | ||||
Stems | unbranched or branched distally, (0.8–)2–8(–10) dm. |
unbranched or branched distally, 1.2–6 dm. |
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Basal leaves | (soon withered); rosulate; petiolate; blade 1- or 2-pinnatifid (with broadly linear to filiform lobes), 2–20 cm, margins sinuate-dentate. |
not rosulate; shortly petiolate; blade lanceolate to oblanceolate, 2–8 cm, margins coarsely dentate to pinnatifid. |
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Cauline leaves | blade linear, 1–10 cm × 1–3 mm, (smaller distally), base auriculate, margins entire. |
blade lanceolate, 1.3–9 cm × 2–20 mm, (smaller distally), base auriculate, margins dentate. |
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Racemes | ebracteate, (lax). |
ebracteate, (lax, with a terminal cluster of dark purple or greenish yellow sterile flowers). |
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Flowers | (markedly zygomorphic); calyx urceolate; sepals greenish yellow or purplish, (abaxial broadly ovate, not keeled, 4–6 × 3–4 mm, lateral ovate-lanceolate, keeled, 4–6 × 1.5–2 mm, adaxial suborbicular to broadly ovate-cordate, forming a bannerlike hood, keeled, 4–6 × 6–8 mm); petals white (with brownish veins), 5–8 mm, blade 1–3 × 0.7–1.2 mm, margins crisped (channeled), claw 4–5 mm, wider than blade; stamens in 3 unequal pairs; filaments: abaxial pair (distinct), 4–5 mm, lateral pair 3–4 mm, adaxial pair (connate), 5–6 mm; anthers: abaxial and lateral pairs fertile, 1.5–2 mm, adaxial pairs sterile, 0.3–0.7 mm; gynophore 0.1–0.4 mm. |
calyx campanulate; sepals (of fertile flowers) dark purple, purplish, or greenish yellow, (oblong to lanceolate), 4–6 mm, (8–14 mm in sterile flowers), not or slightly keeled, (hirsute-hispid in fertile flowers, glabrous in sterile flowers); petals: (abaxial pair spreading and reflexed, adaxial pair parallel and erect), purplish white, lemon-yellow, or yellowish white (with darker midvein), 7–12 mm, blade 2–4 × 1–2 mm, margins crisped, claw 5–8 mm, about as wide as blade; stamens in 3 unequal pairs; filaments: abaxial pair (distinct), 4–6 mm, lateral pair 3–4 mm, adaxial pair (connate their entire length), 6–9 mm; anthers: abaxial and lateral pairs fertile, 1.3–2 mm, adaxial pairs sterile, 0.2–0.7 mm; gynophore 0.4–1 mm. |
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Fruiting pedicels | strongly recurved, 2–5 mm. |
divaricate to ascending, (straight, curved upward, or recurved), 3–8 mm. |
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Fruits | pendent, smooth, straight, flattened, 2.4–5.6 cm × 1.2–1.7 mm; valves each with obscure or somewhat prominent midvein; replum straight; ovules (10–)18–50 per ovary; style 0.8–2 mm; stigma entire. |
ascending or reflexed, smooth, straight or slightly curved, flattened, 3.5–11.4 cm × 1.5–2 mm; valves each with prominent midvein, (glabrate or sparsely to densely hispid, trichomes to 1.7 mm); replum straight; ovules 32–94 per ovary; style 1–2 mm; stigma entire. |
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Seeds | oblong, 1.7–2 × 0.9–1.1 mm; wing 0.2–0.3 mm wide, distal. |
broadly oblong, 1.4–2 × 1–1.3 mm; wing 0.15–0.3 mm wide, continuous. |
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2n | = 28. |
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Streptanthus polygaloides |
Streptanthus insignis |
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Phenology | Flowering May–Jul. | |||||
Habitat | Serpentine substrates in grasslands, openings chaparral, oak and pine woodlands | |||||
Elevation | 200-1900 m (700-6200 ft) | |||||
Distribution |
CA
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CA |
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Discussion | Streptanthus polygaloides is highly variable in plant height, flower color, fruit size, and dissection of basal leaves. Further studies may lead to recognition of infraspecific taxa. The species is distributed in Butte, Calaveras, El Dorado, Fresno, Mariposa, Nevada, Sierra, Tuolumne, and Yuba counties. Streptanthus polygaloides is one of the few nickel hyperaccumulators in the flora area, and it averages 2,430–18,600 µg/g dry weight (R. D. Reeves et al. 1981; A. R. Kruckeberg and Reeves 1995). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 720. | FNA vol. 7, p. 717. | ||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Streptanthus | Brassicaceae > tribe Thelypodieae > Streptanthus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Microsemia polygaloides | |||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 6: 519. (1865) | Jepson: Man. Fl. Pl. Calif., 420. (1925) | ||||
Web links |