Streptanthus polygaloides |
Streptanthus glandulosus |
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milkwort jewelflower |
bristly jewelflower, jewel-flower |
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Habit | Annuals; (sometimes glaucous), glabrous throughout. | Annuals; (usually glaucous distally), often sparsely to densely hirsute proximally, sometimes glabrous throughout. | ||||||||||||||||||||||||||||
Stems | unbranched or branched distally, (0.8–)2–8(–10) dm. |
unbranched or branched throughout, 1.5–12 dm. |
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Basal leaves | (soon withered); rosulate; petiolate; blade 1- or 2-pinnatifid (with broadly linear to filiform lobes), 2–20 cm, margins sinuate-dentate. |
(soon withered); not rosulate; petiolate (petiole winged); blade lanceolate to oblanceolate, 1–15 cm, base narrowed to petiole, margins coarsely dentate to ± lobed. |
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Cauline leaves | blade linear, 1–10 cm × 1–3 mm, (smaller distally), base auriculate, margins entire. |
blade linear-lanceolate to lanceolate or oblanceolate, 1–12 cm × 1–11 mm, (smaller distally), blade auriculate to amplexicaul, margins entire or coarsely dentate, (often entire distally). |
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Racemes | ebracteate, (lax). |
ebracteate, (lax, secund or not, rachis usually straight, rarely flexuous). |
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Flowers | (markedly zygomorphic); calyx urceolate; sepals greenish yellow or purplish, (abaxial broadly ovate, not keeled, 4–6 × 3–4 mm, lateral ovate-lanceolate, keeled, 4–6 × 1.5–2 mm, adaxial suborbicular to broadly ovate-cordate, forming a bannerlike hood, keeled, 4–6 × 6–8 mm); petals white (with brownish veins), 5–8 mm, blade 1–3 × 0.7–1.2 mm, margins crisped (channeled), claw 4–5 mm, wider than blade; stamens in 3 unequal pairs; filaments: abaxial pair (distinct), 4–5 mm, lateral pair 3–4 mm, adaxial pair (connate), 5–6 mm; anthers: abaxial and lateral pairs fertile, 1.5–2 mm, adaxial pairs sterile, 0.3–0.7 mm; gynophore 0.1–0.4 mm. |
(zygomorphic); calyx urceolate; sepals white, greenish white, cream, yellow, rose, purple, lilac, maroon, reddish purple, or purple-black, (lanceolate to broadly ovate), (3–)5–10(–13) mm, not keeled, (glabrous or sparsely hirsute); petals lavender, purple, or white (sometimes with brown or purple veins), 7–17 mm, blade 2–7 × 1–3 mm, (subequal or adaxial pair distinctly longer, more recurved), margins crisped, claw 5–13 mm, wider than blade; stamens in 3 unequal pairs; filaments: abaxial pair (distinct), 4–9 mm, lateral pair 2.5–7 mm, adaxial pair (exserted, connate 2/3 their length), 5–13 mm; anthers: abaxial and lateral pairs fertile, 1–2.5 mm, adaxial pair sterile, 0.3–1 mm; gynophore 0.2–1.5 mm. |
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Fruiting pedicels | strongly recurved, 2–5 mm. |
ascending to divaricate or spreading, (straight or recurved), 2–32 mm, (glabrous or pubescent). |
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Fruits | pendent, smooth, straight, flattened, 2.4–5.6 cm × 1.2–1.7 mm; valves each with obscure or somewhat prominent midvein; replum straight; ovules (10–)18–50 per ovary; style 0.8–2 mm; stigma entire. |
ascending to divaricate or spreading, smooth, straight, curved upward, or arcuate, flattened, 3–11 cm × 1.5–2.5 mm; valves each with prominent midvein, (glabrous or sparsely to moderately hirsute); replum straight; ovules 22–70 per ovary; style 0.1–2.5 mm; stigma subentire. |
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Seeds | oblong, 1.7–2 × 0.9–1.1 mm; wing 0.2–0.3 mm wide, distal. |
ovoid to oblong, 1.5–2.1 × 1–1.5 mm; wing 0.1–0.5 mm wide, continuous. |
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2n | = 28. |
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Streptanthus polygaloides |
Streptanthus glandulosus |
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Phenology | Flowering May–Jul. | |||||||||||||||||||||||||||||
Habitat | Serpentine substrates in grasslands, openings chaparral, oak and pine woodlands | |||||||||||||||||||||||||||||
Elevation | 200-1900 m (700-6200 ft) | |||||||||||||||||||||||||||||
Distribution |
CA
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CA; OR
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Discussion | Streptanthus polygaloides is highly variable in plant height, flower color, fruit size, and dissection of basal leaves. Further studies may lead to recognition of infraspecific taxa. The species is distributed in Butte, Calaveras, El Dorado, Fresno, Mariposa, Nevada, Sierra, Tuolumne, and Yuba counties. Streptanthus polygaloides is one of the few nickel hyperaccumulators in the flora area, and it averages 2,430–18,600 µg/g dry weight (R. D. Reeves et al. 1981; A. R. Kruckeberg and Reeves 1995). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 8 (8 in the flora). Except for the widespread subsp. glandulosus, the subspecies are highly restricted and six are of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 720. | FNA vol. 7, p. 712. | ||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Streptanthus | Brassicaceae > tribe Thelypodieae > Streptanthus | ||||||||||||||||||||||||||||
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Synonyms | Microsemia polygaloides | Erysimum glandulosum, Euklisia glandulosa | ||||||||||||||||||||||||||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 6: 519. (1865) | Hooker: Icon. Pl. 1: plate 40. (1836) | ||||||||||||||||||||||||||||
Web links |