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milkwort jewelflower

bristly jewelflower, jewel-flower

Habit Annuals; (sometimes glaucous), glabrous throughout. Annuals; (usually glaucous distally), often sparsely to densely hirsute proximally, sometimes glabrous throughout.
Stems

unbranched or branched distally, (0.8–)2–8(–10) dm.

unbranched or branched throughout, 1.5–12 dm.

Basal leaves

(soon withered); rosulate; petiolate;

blade 1- or 2-pinnatifid (with broadly linear to filiform lobes), 2–20 cm, margins sinuate-dentate.

(soon withered);

not rosulate; petiolate (petiole winged);

blade lanceolate to oblanceolate, 1–15 cm, base narrowed to petiole, margins coarsely dentate to ± lobed.

Cauline leaves

blade linear, 1–10 cm × 1–3 mm, (smaller distally), base auriculate, margins entire.

blade linear-lanceolate to lanceolate or oblanceolate, 1–12 cm × 1–11 mm, (smaller distally), blade auriculate to amplexicaul, margins entire or coarsely dentate, (often entire distally).

Racemes

ebracteate, (lax).

ebracteate, (lax, secund or not, rachis usually straight, rarely flexuous).

Flowers

(markedly zygomorphic);

calyx urceolate;

sepals greenish yellow or purplish, (abaxial broadly ovate, not keeled, 4–6 × 3–4 mm, lateral ovate-lanceolate, keeled, 4–6 × 1.5–2 mm, adaxial suborbicular to broadly ovate-cordate, forming a bannerlike hood, keeled, 4–6 × 6–8 mm);

petals white (with brownish veins), 5–8 mm, blade 1–3 × 0.7–1.2 mm, margins crisped (channeled), claw 4–5 mm, wider than blade;

stamens in 3 unequal pairs;

filaments: abaxial pair (distinct), 4–5 mm, lateral pair 3–4 mm, adaxial pair (connate), 5–6 mm;

anthers: abaxial and lateral pairs fertile, 1.5–2 mm, adaxial pairs sterile, 0.3–0.7 mm;

gynophore 0.1–0.4 mm.

(zygomorphic);

calyx urceolate;

sepals white, greenish white, cream, yellow, rose, purple, lilac, maroon, reddish purple, or purple-black, (lanceolate to broadly ovate), (3–)5–10(–13) mm, not keeled, (glabrous or sparsely hirsute);

petals lavender, purple, or white (sometimes with brown or purple veins), 7–17 mm, blade 2–7 × 1–3 mm, (subequal or adaxial pair distinctly longer, more recurved), margins crisped, claw 5–13 mm, wider than blade;

stamens in 3 unequal pairs;

filaments: abaxial pair (distinct), 4–9 mm, lateral pair 2.5–7 mm, adaxial pair (exserted, connate 2/3 their length), 5–13 mm;

anthers: abaxial and lateral pairs fertile, 1–2.5 mm, adaxial pair sterile, 0.3–1 mm;

gynophore 0.2–1.5 mm.

Fruiting pedicels

strongly recurved, 2–5 mm.

ascending to divaricate or spreading, (straight or recurved), 2–32 mm, (glabrous or pubescent).

Fruits

pendent, smooth, straight, flattened, 2.4–5.6 cm × 1.2–1.7 mm;

valves each with obscure or somewhat prominent midvein;

replum straight;

ovules (10–)18–50 per ovary;

style 0.8–2 mm;

stigma entire.

ascending to divaricate or spreading, smooth, straight, curved upward, or arcuate, flattened, 3–11 cm × 1.5–2.5 mm;

valves each with prominent midvein, (glabrous or sparsely to moderately hirsute);

replum straight;

ovules 22–70 per ovary;

style 0.1–2.5 mm;

stigma subentire.

Seeds

oblong, 1.7–2 × 0.9–1.1 mm;

wing 0.2–0.3 mm wide, distal.

ovoid to oblong, 1.5–2.1 × 1–1.5 mm;

wing 0.1–0.5 mm wide, continuous.

2n

= 28.

Streptanthus polygaloides

Streptanthus glandulosus

Phenology Flowering May–Jul.
Habitat Serpentine substrates in grasslands, openings chaparral, oak and pine woodlands
Elevation 200-1900 m (700-6200 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Streptanthus polygaloides is highly variable in plant height, flower color, fruit size, and dissection of basal leaves. Further studies may lead to recognition of infraspecific taxa. The species is distributed in Butte, Calaveras, El Dorado, Fresno, Mariposa, Nevada, Sierra, Tuolumne, and Yuba counties.

Streptanthus polygaloides is one of the few nickel hyperaccumulators in the flora area, and it averages 2,430–18,600 µg/g dry weight (R. D. Reeves et al. 1981; A. R. Kruckeberg and Reeves 1995).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 8 (8 in the flora).

Except for the widespread subsp. glandulosus, the subspecies are highly restricted and six are of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems proximally and cauline leaf blade surfaces glabrous; sepals purple-black; rachises flexuous; fruiting pedicels 10-32 mm.
subsp. niger
1. Stems proximally and cauline leaf blade surfaces usually sparsely to densely hirsute (cauline leaves glabrous in subsp. albidus); sepals white, greenish white, cream, yellow, lilac, lavender, rose, purple, maroon, or reddish purple; rachises straight; fruiting pedicels 2-15 mm
→ 2
2. Racemes not secund
→ 3
2. Racemes secund
→ 4
3. Sepals reddish purple, dark maroon, or lilac-lavender; San Luis Obispo County north into Lake, Mendocino, Santa Clara counties.
subsp. glandulosus
3. Sepals white, greenish white, or pale yellow; Santa Clara County.
subsp. albidus
4. Sepals lavender to rose or purple
→ 5
4. Sepals greenish white, white, cream, or pale yellow
→ 6
5. Fruiting pedicels 5-15 mm; fruits spreading to reflexed, arcuate; filaments: adaxial pair 7-10 mm; Sonoma County.
subsp. hoffmanii
5. Fruiting pedicels 2-5 mm; fruits divaricate to ascending, curved upward; filaments: adaxial pair 6-8 mm; Marin County.
subsp. pulchellus
6. Petals 7-8 mm; filaments: adaxial pair 5-6 mm; fruits divaricate-ascending, straight or curved inwards; Josephine County, Oregon.
subsp. josephinensis
6. Petals 10-17 mm; filaments: adaxial pair 7-10 mm; fruits spreading to reflexed, arcuate; Marin, Napa, Sonoma counties, California
→ 7
7. Sepals greenish white with lavender or purplish base; petals with purple veins; Marin, Napa counties.
subsp. secundus
7. Sepals cream or pale yellow, without purplish or lavender base; petals without purple veins; Sonoma County.
subsp. sonomensis
Source FNA vol. 7, p. 720. FNA vol. 7, p. 712.
Parent taxa Brassicaceae > tribe Thelypodieae > Streptanthus Brassicaceae > tribe Thelypodieae > Streptanthus
Sibling taxa
S. barbatus, S. barbiger, S. batrachopus, S. bernardinus, S. brachiatus, S. bracteatus, S. breweri, S. callistus, S. campestris, S. carinatus, S. cordatus, S. cutleri, S. diversifolius, S. drepanoides, S. farnsworthianus, S. fenestratus, S. glandulosus, S. gracilis, S. hesperidis, S. hispidus, S. howellii, S. hyacinthoides, S. insignis, S. longisiliquus, S. maculatus, S. morrisonii, S. oblanceolatus, S. oliganthus, S. petiolaris, S. platycarpus, S. squamiformis, S. tortuosus, S. vernalis, S. vimineus
S. barbatus, S. barbiger, S. batrachopus, S. bernardinus, S. brachiatus, S. bracteatus, S. breweri, S. callistus, S. campestris, S. carinatus, S. cordatus, S. cutleri, S. diversifolius, S. drepanoides, S. farnsworthianus, S. fenestratus, S. gracilis, S. hesperidis, S. hispidus, S. howellii, S. hyacinthoides, S. insignis, S. longisiliquus, S. maculatus, S. morrisonii, S. oblanceolatus, S. oliganthus, S. petiolaris, S. platycarpus, S. polygaloides, S. squamiformis, S. tortuosus, S. vernalis, S. vimineus
Subordinate taxa
S. glandulosus subsp. albidus, S. glandulosus subsp. glandulosus, S. glandulosus subsp. hoffmanii, S. glandulosus subsp. josephinensis, S. glandulosus subsp. niger, S. glandulosus subsp. pulchellus, S. glandulosus subsp. secundus, S. glandulosus subsp. sonomensis
Synonyms Microsemia polygaloides Erysimum glandulosum, Euklisia glandulosa
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 519. (1865) Hooker: Icon. Pl. 1: plate 40. (1836)
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