Streptanthus polygaloides |
Streptanthus cordatus |
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milkwort jewelflower |
heart-leaf twist-flower, heartleaf jewelflower |
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Habit | Annuals; (sometimes glaucous), glabrous throughout. | Perennials; (caudex simple or few-branched); (glaucous), usually glabrous, (petioles of basal leaves pubescent, sometimes sepals). | ||||
Stems | unbranched or branched distally, (0.8–)2–8(–10) dm. |
unbranched or branched, (1–)3–9(–11) dm. |
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Basal leaves | (soon withered); rosulate; petiolate; blade 1- or 2-pinnatifid (with broadly linear to filiform lobes), 2–20 cm, margins sinuate-dentate. |
rosulate; petiolate (petioles broadly winged, setose-ciliate); blade usually spatulate to obovate, rarely narrowly oblanceolate, 1.5–8(–11) cm, margins dentate (at least distally). |
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Cauline leaves | blade linear, 1–10 cm × 1–3 mm, (smaller distally), base auriculate, margins entire. |
blade broadly oblong to ovate, suborbicular, or lanceolate, 2–9 cm × 7–45(–60) mm, (smaller distally), base auriculate to amplexicaul, margins entire or toothed, (apex rounded, obtuse, or acuminate to acute). |
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Racemes | ebracteate, (lax). |
ebracteate, (lax). |
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Flowers | (markedly zygomorphic); calyx urceolate; sepals greenish yellow or purplish, (abaxial broadly ovate, not keeled, 4–6 × 3–4 mm, lateral ovate-lanceolate, keeled, 4–6 × 1.5–2 mm, adaxial suborbicular to broadly ovate-cordate, forming a bannerlike hood, keeled, 4–6 × 6–8 mm); petals white (with brownish veins), 5–8 mm, blade 1–3 × 0.7–1.2 mm, margins crisped (channeled), claw 4–5 mm, wider than blade; stamens in 3 unequal pairs; filaments: abaxial pair (distinct), 4–5 mm, lateral pair 3–4 mm, adaxial pair (connate), 5–6 mm; anthers: abaxial and lateral pairs fertile, 1.5–2 mm, adaxial pairs sterile, 0.3–0.7 mm; gynophore 0.1–0.4 mm. |
calyx campanulate; sepals greenish brown to purple, (broadly oblong), 5–12 mm, (lateral pair subsaccate basally), not keeled, (glabrous or subapically bristly); petals purple to brownish, (claw purplish), 9–15 mm, blade 2–6 × 0.7–1 mm, (recurved or not), margins not crisped, claw 7–10 mm, wider than blade; stamens in 3 unequal pairs; filaments (distinct): abaxial pair 5–7 mm, lateral pair 4–5 mm, adaxial pair 7.5–10 mm; anthers (all) fertile, 2.5–5 mm; gynophore 0.5–1.5 mm. |
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Fruiting pedicels | strongly recurved, 2–5 mm. |
divaricate-ascending, (straight), 3–11(–14) mm. |
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Fruits | pendent, smooth, straight, flattened, 2.4–5.6 cm × 1.2–1.7 mm; valves each with obscure or somewhat prominent midvein; replum straight; ovules (10–)18–50 per ovary; style 0.8–2 mm; stigma entire. |
ascending to divaricate-ascending, smooth, straight, flattened, 5–10.5(–14.5) cm × (2.5–)3–6(–7) mm; valves each with prominent midvein; replum straight; ovules 20–38(–46) per ovary; style 0.2–3 mm; stigma subentire to strongly 2-lobed. |
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Seeds | oblong, 1.7–2 × 0.9–1.1 mm; wing 0.2–0.3 mm wide, distal. |
broadly oblong to suborbicular, 2.5–5 × 2.2–5 mm; wing 0.1–0.9 mm wide, continuous. |
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2n | = 28. |
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Streptanthus polygaloides |
Streptanthus cordatus |
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Phenology | Flowering May–Jul. | |||||
Habitat | Serpentine substrates in grasslands, openings chaparral, oak and pine woodlands | |||||
Elevation | 200-1900 m (700-6200 ft) | |||||
Distribution |
CA
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AZ; CA; CO; ID; NM; NV; OR; UT; WY
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Discussion | Streptanthus polygaloides is highly variable in plant height, flower color, fruit size, and dissection of basal leaves. Further studies may lead to recognition of infraspecific taxa. The species is distributed in Butte, Calaveras, El Dorado, Fresno, Mariposa, Nevada, Sierra, Tuolumne, and Yuba counties. Streptanthus polygaloides is one of the few nickel hyperaccumulators in the flora area, and it averages 2,430–18,600 µg/g dry weight (R. D. Reeves et al. 1981; A. R. Kruckeberg and Reeves 1995). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Streptanthus cordatus is the most widespread species in the genus. It is highly variable in the shape and size of basal and cauline leaves, flower size, pedicel length, fruit length and width, style length, stigma lobing, seed shape and size, and width of the seed wing. The variation is continuous in almost every character, especially fruit width. R. E. Buck et al. (1993) indicated that in var. cordatus the fruits are more than 5 mm wide and in the Californian var. duranii they are less than 3 mm wide. One sporadically finds throughout the range of var. cordatus plants with mature fruits 2.5–3 mm wide. Such narrow-fruited forms are found in central Nevada (Beatley 6037, DS), Oregon (Howell 28719, CAS), Utah (Goodman 1877, MO), and Wyoming (Rollins & Muñoz 2876, GH). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 720. | FNA vol. 7, p. 708. | ||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Streptanthus | Brassicaceae > tribe Thelypodieae > Streptanthus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Microsemia polygaloides | Cartiera cordata, Erysimum cordatum, Euklisia cordata | ||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 6: 519. (1865) | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 77. (1838) | ||||
Web links |