Streptanthus campestris |
Streptanthus tortuosus |
|
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southern jewel flower |
jewelweed, mountain jewel-flower, shield plant, shieldleaf |
|
Habit | Perennials; (short-lived, caudex not woody); usually glabrous, (basal leaf blade margins pubescent, sometimes sepals). | Biennials or perennials; (short-lived); (glaucous), glabrous throughout. |
Stems | unbranched or branched, (few, glaucous), 6–15(–18) dm. |
unbranched or branched (several) basally and distally, (0.5–)1.5–12(–15) dm. |
Basal leaves | often rosulate; petiolate; blade (fleshy), oblanceolate to obovate, 3.5–21 cm, margins dentate, (bristly ciliate throughout or only teeth and petiole ciliate). |
(soon withered); subrosulate; petiolate; blade broadly ovate, obovate or oblong, 1.5–6 cm, margins entire or repand, or dentate apically. |
Cauline leaves | blade lanceolate to narrowly ovate, 3.5–11(–15) cm × 6–14 mm (smaller distally), base auriculate to amplexicaul, margins usually entire or undulate, rarely dentate. |
blade oblong to obovate or suborbicular, or (distally) orbicular or oblong-ovate, (0.7–)1.5–6(–9) cm × (4–)10–45 mm, base auriculate to amplexicaul (amplexicaul distally), margins entire or repand. |
Racemes | ebracteate, (with densely clustered buds, later lax). |
bracteate below or between proximalmost 1 or 2 flowers, (lax to compact, secund or not). |
Flowers | calyx campanulate; sepals (suberect), purple, (broadly ovate or oblong), 7–10 mm, not keeled, (apically bristly or not); petals light purple (with pale yellow claw), 9–12 mm, blade 2–3.5 × 0.5–1 mm, margins not crisped, claw 6–9 mm, wider than blade; stamens tetradynamous; filaments: median pairs (distinct), 6–8 mm, lateral pair 4–6 mm; anthers (all) fertile, 3–4 mm; gynophore 0.5–1.5 mm. |
calyx urceolate; sepals purplish, gray-green, or yellowish, 6–10(–13) mm, keeled or not, (apex recurved); petals purplish or yellowish white (usually with purple veins), (6–)8–14 mm, blade 2–5 × 1–2.5 mm, margins not crisped, claw 4–10 mm, as wide as or wider than blade, (apex reflexed); stamens in 3 unequal pairs; filaments (distinct): abaxial pair (2.5–)4–7 mm, lateral pair (1.5–)3–5 mm, adaxial pair (5–)7–11 mm; anthers (all) fertile or adaxial pair with reduced fertility and shorter, (1.5–)2.5–4.5(–6) mm; gynophore 0.2–1 mm. |
Fruiting pedicels | divaricate-ascending, (straight), 5–18 mm. |
divaricate to ascending, (2–)3–12(–17) mm, (expanded at receptacle). |
Fruits | spreading to ascending, smooth, slightly curved to straight, flattened, 6–14 cm × 2–3.5 mm; valves each with obscure midvein; replum straight; ovules 50–102 per ovary; style 1–3 mm; stigma 2-lobed. |
arcuate-spreading to pendent, torulose or smooth, straight, flattened, (3–)4–13(–16) cm × 1.5–2.5(–3) mm; valves each with obscure or somewhat prominent midvein; replum straight; ovules 26–76(–110) per ovary; style 0.4–0.7 mm; stigma entire. |
Seeds | oblong, 2–3 × 1.4–2 mm; wing 0.1–0.2 mm wide at apex. |
broadly oblong to ovoid or orbicular, 1.5–2.5 × 0.8–1.7 mm; wing 0.1–0.5 mm wide, continuous or on distal 1/2. |
2n | = 28. |
|
Streptanthus campestris |
Streptanthus tortuosus |
|
Phenology | Flowering May–Jun. | Flowering Apr–Sep. |
Habitat | Rocky openings in chaparral, open conifer forests, openings and after fires in chaparral-oak woodlands | Rocky open slopes, open woodlands, montane forests, alpine areas |
Elevation | 900-2300 m (3000-7500 ft) | 200-4100 m (700-13500 ft) |
Distribution |
CA; Mexico (Baja California)
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CA; NV; OR
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Discussion | Of conservation concern. Streptanthus campestris is distributed in California in Riverside, San Bernardino, San Diego, Santa Barbara, and Ventura counties, and in Baja California in Sierra San Pedro Mártir and Sierra Juárez. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Streptanthus tortuosus is highly polymorphic, forming distinct local races. Variation in duration, stature, branching, development of the sepal keel, petal color, and bract size are not correlated with habitat or geography. Some authors (e.g., R. C. Rollins 1993; R. E. Buck et al. 1993) divided it into five varieties based on variations in those characters, but the variation is continuous and the purported differences do not hold. For example, those authors and N. H. Holmgren (2005b), recognized the highly branched, shorter plants of the alpine Sierra Nevada as var. orbiculatus, but intergradation downslope is completely clinal. The most distinctive variant (S. foliosus) is restricted to the central Sierra Nevada, where plants with large, brittle, subachlorophyllus bracts predominate. However, that condition reappears sporadically in the Klamath Ranges and elsewhere. Annual plants have never been documented in S. tortuosus, contrary to the claim by some authors (e.g., Rollins; Buck et al.; Holmgren); the alleged “annuals” produce rosettes that invariably overwinter before flowering. Without thorough molecular, experimental, and cytological study of this complex, it is impractical to recognize ill-defined varieties that represent only a minor fraction of the overall variation in the species. R. C. Rollins (1993) placed Streptanthus foliosus in the synonymy of S. diversifolius, but its type clearly belongs to S. tortuosus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 707. | FNA vol. 7, p. 722. |
Parent taxa | Brassicaceae > tribe Thelypodieae > Streptanthus | Brassicaceae > tribe Thelypodieae > Streptanthus |
Sibling taxa | ||
Synonyms | Disaccanthus tortuosus, Erysimum tortuosum, Pleiocardia foliosa, Pleiocardia orbiculata, Pleiocardia suffrutescens, Pleiocardia tortuosa, S. foliosus, S. orbiculatus, S. sanhedrensis, S. suffrutescens, S. tortuosus var. flavescens, S. tortuosus var. oblongus, S. tortuosus var. optatus, S. tortuosus var. orbiculatus, S. tortuosus var. pallidus, S. tortuosus var. suffrutescens | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 25: 125. (1890) | Kellogg: Proc. Calif. Acad. Sci. 2: 152, plate 46. (1863) |
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