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Trecul's toothleaf

Mojave stillingia, Mojave toothleaf, tooth leaf

Habit Herbs, perennial, with woody taproot. Herbs, perennial, with thick taproot.
Stems

fascicled, spreading, branching throughout, 1–4.5 dm.

fascicled, erect, branching scattered proximally and crowded distally, (1.4–)2–3.5(–4) dm.

Leaves

alternate;

stipules linear, to 1 mm;

petiole absent;

blade usually obovate-spatulate, rarely narrowly elliptic, 1–-4 × 0.5–1.5 cm, base cuneate, margins coarsely spinulose-dentate, teeth without prominent blackened tips, not incurved, apex rounded to obtuse;

midrib prominent, secondary veins obscure.

alternate;

stipules absent;

petiole absent;

blade linear, 2–4(–6) × 0.1–0.3(–0.4) cm, base acute, margins entire or remotely denticulate, teeth without prominent blackened tips, not incurved, apex usually acute, rarely acuminate;

venation not prominent.

Inflorescences

pedunculate, 2.5–-5 cm;

staminate flowers ± widely spaced, 1 per node;

pistillate flowers 3–4, widely spaced;

bracts ovate, 1 mm, apex mucronate, glands patelliform, ± sessile, 0.7 mm diam.

sessile or rarely short-pedunculate, 6–-7 cm;

staminate flowers crowded distally, 1 per node;

pistillate flowers 3–5, crowded;

bracts broadly ovate, to 1.5 mm, apex mucronulate or acuminate, glands patelliform, sessile, to 1.3 mm diam.

Staminate flowers

calyx 1 mm.

calyx to 1 mm.

Pistillate flowers

sepals fugacious, 3, minute;

styles connate only at base, 1.5 mm.

sepals 0;

styles connate only at base, to 4 mm.

Capsules

ovoid-pyriform, 4 × 5 mm, deeply 3-lobed;

lobes of gynobase to 1 mm;

columella persistent.

oblate, 3 × 4 mm, deeply 3-lobed;

lobes of gynobase 1.5–2 mm;

columella persistent.

Seeds

gray to black, ellipsoid, 2.5 × 2 mm, smooth;

caruncle minute.

brown, often mottled, ovoid, 2.3 × 1.3 mm, smooth;

caruncle minute.

Stillingia treculiana

Stillingia paucidentata

Phenology Flowering Mar–May; fruiting (Mar–)Apr–Jul(–Dec). Flowering Mar–May; fruiting May–Jun.
Habitat Calcareous sandy soils, gravelly soils, and uplands. Sandy flats, dry slopes.
Elevation 0–1000 m. (0–3300 ft.) 0–1500 m. (0–4900 ft.)
Distribution
from FNA
TX; Mexico (Coahuila, Nuevo León, Tamaulipas)
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Stillingia treculiana is known in the flora area from the western Edwards Plateau south to the lower Rio Grande valley.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Stillingia paucidentata is widespread in the Mojave Desert and extends into the Sonoran Desert in central Riverside County. It was reported from Arizona by T. H. Kearney and R. H. Peebles (1942, 1960) solely on the basis of the type (Palmer 517), purportedly collected in 1876 in the “Colorado Valley, near mouth of Williams River.” R. McVaugh (1943b) and McVaugh and Kearney (1943) have cast doubt on whether a number of Palmer collections with labels indicating 1876 were actually made in Arizona; they did not discuss Palmer 517 specifically. There appear to be no other specimens or reports of this species from Arizona. Because S. paucidentata is known from numerous collections in eastern San Bernardino County, California, its presence in bordering areas of Arizona cannot be completely discounted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 237. FNA vol. 12, p. 235.
Parent taxa Euphorbiaceae > Stillingia Euphorbiaceae > Stillingia
Sibling taxa
S. aquatica, S. linearifolia, S. paucidentata, S. spinulosa, S. sylvatica, S. texana
S. aquatica, S. linearifolia, S. spinulosa, S. sylvatica, S. texana, S. treculiana
Synonyms Gymnanthes treculiana
Name authority (Müller Arg.) I. M. Johnston: Contr. Gray Herb. 68: 91. (1923) S. Watson: Proc. Amer. Acad. Arts 14: 298. (1879)
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