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blunt-sepal starwort, obtuse starwort, Rocky Mountain chickweed, Rocky Mountain starwort

chickweed, starwort, stellaire, stitchwort

Habit Plants perennial, creeping, often matted but not forming cushions, rhizomatous. Plants annual, winter annual, or perennial.
Taproots

usually slender, perennial taxa often rhizomatous, rooting at nodes.

Stems

prostrate, branched, 4-sided, 3–23 cm, internodes equaling or longer than leaves, glabrous, rarely pilose.

prostrate to ascending or erect, simple or branched, terete or 4-angled.

Leaves

sessile or short-petiolate;

blade broadly ovate to elliptic, 0.2–1.2 cm × 0.9–7 mm, base round or cuneate, margins entire, apex acute, shiny, glabrous or ciliate near base.

sometimes connate basally into sheath, often sessile;

blade 1-veined, linear or lanceolate to ovate or deltate, succulent (S. crassifolia [gemmae], S. fontinalis, S. humifusa, and S. irrigua) or not, apex acute or obtuse.

Inflorescences

with flowers solitary, axillary;

bracts absent.

terminal, open cymes, rarely axillary (S. alsine, S. americana) or umbellate (S. umbellata), or terminal or axillary solitary flowers;

bracts paired (1 in S. dicranoides), foliaceous, scarious and reduced, or absent.

Pedicels

spreading, 3–12 mm, glabrous.

erect, sometimes reflexed in fruit, glabrous or pubescent, not glandular.

Flowers

1.5–2 mm diam.;

sepals 4–5, veins obscure, midrib sometimes apparent, ± ovate, 1.5–3.5 mm, margins narrow, scarious, apex ± obtuse, glabrous;

petals absent;

stamens 10 or fewer;

styles 3(–4), curled, shorter than 0.5 mm.

usually bisexual (S. dicranoides unisexual);

perianth and androecium hypogynous or weakly perigynous;

hypanthium cup- or disc-shaped;

sepals (4–)5, distinct, green, occasionally purple tinged (S. irrigua) or red proximally (S. pallida), lanceolate to ovate-triangular, 2–12 mm, herbaceous (rarely coriaceous), margins often white, scarious, apex acute, acuminate, or obtuse, not hooded;

petals (1–)5 or absent, white (sometimes translucent in S. borealis), not clawed, blade apex 2-fid usually for 2/3–4/5 its length (S. holostea occasionally laciniate);

nectaries at base of filaments opposite sepals usually present, disc sometimes prominent;

stamens (1–)5 or 10 or absent, arising from nectariferous disc (prominent in S. dicranoides and S. irrigua) at ovary base;

filaments distinct;

staminodes absent;

styles [2–]3(–5), capitate to clavate, 0.2–7 mm, glabrous proximally;

stigmas [2–]3(–5), terminal or subterminal, papillate (30x).

Capsules

green to pale straw colored, translucent, globose to broadly ovoid, 2.3–3.5 mm, 1.9–2 times as long as sepals, apex obtuse, opening by 6 valves;

carpophore absent.

globose to conic, opening by 3 or 6, occasionally 4, 8, or 10 ascending to recurved valves;

carpophore present or absent.

Seeds

grayish black, broadly elliptic, 0.5–0.7 mm diam., finely reticulate.

(1–)3–20+, yellow-brown to dark brown, globose to ellipsoid, laterally compressed, rarely shiny, papillate or rugose, rarely smooth, marginal wing absent, appendage absent.

x

= 10, 11, 12, 13, 15.

2n

= 26, 52, ca. 65, ca. 78.

Stellaria obtusa

Stellaria

Phenology Flowering late spring–summer.
Habitat Moist areas in woods, shaded edges of creeks, talus slopes
Elevation 300-3400 m (1000-11200 ft)
Distribution
from FNA
CA; CO; ID; MT; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from USDA
Worldwide; mainly north-temperate regions
[BONAP county map]
Discussion

Species ca. 120 (29 in the flora).

I have not attempted to present an infrageneric classification for Stellaria. Although several species complexes can be identified within the genus (e.g., S. media, S. longipes, S. calycantha), there are significant problems with accepting the most recent scheme (F. Pax and K. Hoffmann 1934c). Stellaria jamesiana is now placed in Pseudostellaria and S. aquatica in Myosoton. As well, S. dicranoides and S. fontinalis probably should be placed in other genera (see comments under those species). Additional study is warranted to determine both the generic circumscription of Stellaria and species relationships within the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Mid-stem and proximal leaves distinctly petiolate, broadly lanceolate to ovate or deltate
→ 2
1. Mid-stem and proximal leaves sessile to shortly petiolate; blades elliptic, ovate, lanceolate, linear, or linear-lanceolate
→ 8
2. Leaf blade bases cordate to truncate (rarely abruptly rounded)
S. cuspidata
2. Leaf blade bases round or cuneate
→ 3
3. Flowers large, usually 10 mm or more diam.; petals usually exceeding sepals
→ 4
3. Petals equaling or shorter than sepals, or absent; flowers less than 10 mm diam
→ 5
4. Sepals obtuse to acute, 3.5-6 mm
S. pubera
4. Sepals acuminate, (5-)7-10(-12) mm
S. corei
5. Flowers solitary in axils of mid and distal stem leaves
S. obtusa
5. Flowers several in foliaceous terminal cymes
→ 6
6. Sepals 5-6.5 mm; stamens 8-10; seeds 1.1-1.7 mm; tubercles taller than broad, apex acute
S. neglecta
6. Sepals usually 3-5(-6) mm; stamens 1-5(-8); seeds 0.4-1.3 mm; tubercles usually broader than tall, apex usually obtuse
→ 7
7. Stamens 3-5(-8); sepals 4.5-5(-6) mm; seeds 0.9-1.3 mm diam.; petals usually present; plants usually green
S. media
7. Stamens 1-3 or absent; sepals 3-4 mm; seeds mostly 0.5-0.9 mm diam.; petals usually absent; plants usually yellowish green
S. pallida
8. Inflorescences, or flowers when solitary, in axils of foliage leaves on mid or distal stem
→ 9
8. Inflorescences with most flowers terminal, either several in bracteate inflorescence or solitary on long-ascending pedicels
→ 18
9. Petals equaling or longer than sepals
→ 10
9. Petals shorter than sepals, or absent
→ 13
10. Sepals narrowly lanceolate-triangular, prominently 3-veined, margins straight
S. crassifolia
10. Sepals lanceolate to ovate-lanceolate, 1- or obscurely 3-veined, margins convex
→ 11
11. Seeds prominently papillate, papillae taller than broad; sepals pubescent
S. parva
11. Seeds smooth, rugose, or shallowly tuberculate, not papillate; sepals glabrous
→ 12
12. Leaf blades succulent, elliptic to elliptic-lanceolate, 4-15 × 1-5 mm
S. humifusa
12. Leaf blades not succulent, linear to linear-lanceolate, 27-35 × 2-3 mm
S. porsildii
13. Plants forming dense cushions, mid and distal stem internodes shorter than leaves
→ 14
13. Plants creeping to straggling or ascending, sometimes forming mats, but not forming cushions, internodes equaling or longer than leaves
→ 15
14. Stamens 5; petals present, deeply divided into 2 narrowly elliptic lobes
S. irrigua
14. Stamens 10; petals absent
S. dicranoides
15. Leaf blades narrowly elliptic to lanceolate, oblanceolate, or linear
→ 16
15. Leaf blades broadly elliptic to ovate
→ 17
16. Flowers solitary in distal leaf axils
S. fontinalis
16. Flowers in axillary inflorescences in mid and distal leaf axils
S. alsine
17. Sepals obtuse at apex with ± obscure veins; styles shorter than 0.5 mm, curled; capsules globose to broadly ovoid
S. obtusa
17. Sepals acute to acuminate at apex with 3 prominent veins; styles ca. 1 mm, spreading to ascending; capsules ovoid to ovoid-ellipsoid
S. crispa
18. Bracts scarious or with scarious margins
→ 19
18. Bracts or subtending leaf blades without scarious margins
→ 26
19. Capsules ca. equaling or shorter than sepals
→ 20
19. Capsules longer than sepals
→ 22
20. Plants annual
S. nitens
20. Plants perennial
→ 21
21. Plants compact; stems ascending, 3-10(-20) cm; petals equaling or shorter than sepals
S. alaskana
21. Plants coarse; stems straggling with erect branches, 20-60 cm; petals 1.5-2 times as long as sepals
S. palustris
22. Inflorescences subumbellate; petals absent
S. umbellata
22. Inflorescences cymose or flowers solitary; petals present
→ 23
23. Plants delicate, creeping, often forming mats; flowers solitary and axillary or in small, few-flowered, leafy cymes; leaf blades variable in shape, midribobscure
S. crassifolia
23. Plants not with the above combination of characters; leaf blades with prominent midrib
→ 24
24. Leaf blades linear-elliptic, broadest at middle or distally; angles of stems and/or margins of leaf blades minutely papillate-scabrid (30×)
S. longifolia
24. Leaf blades lanceolate to linear-lanceolate, broadest proximally; angles of stems and/or margins of leaf blades not papillate-scabrid (soft hairs of cilia may be present)
→ 25
25. Sepals narrowly triangular or lanceolate, margins straight, veins 3, forming prominent ridges; seeds coarsely rugose in concentric rings; plants coarse; stems ascending, straggling; inflorescences muchbranched, many-flowered
S. graminea
25. Sepals lanceolate to ovate-lanceolate, margins convex, veins 1-3, not forming ridges; seeds shallowly tuberculate to smooth; stems erect to straggling; flowers solitary or inflorescences few-flowered
S. longipes
26. Flowers 20-30 mm diam.; petals 8-14 mm, longer than sepals; leaf blades narrowly lanceolate, 4-8 cm, apex acuminate
S. holostea
26. Plants not having the above combination of characters
→ 27
27. Leaves widest at or above middle
→ 28
27. Leaves widest towards base
→ 31
28. Plants glandular-puberulent; pedicels arcuate, pushing capsule into substrate; seeds 3-6, ca. 2.5 mm diam
S. americana
28. Plants eglandular; other characters not as above
→ 29
29. Sepals narrowly lanceolate-triangular; plants creeping, forming mats usually shorter than 5 cm
S. crassifolia
29. Sepals ovate to ovate-triangular; plants with diffusely branched or ascending flowering stems taller than 5 cm
→ 30
30. Petals 4-8 mm, conspicuous, white; sepals 3.5-6 mm
S. pubera
30. Petals shorter than 3 mm, inconspicuous, white or translucent; sepals2-3(-4) mm
S. borealis
31. Plants straggling to scandent; stems to 60 cm; leaf blades ovate to ovate-lanceolate, 10-45 × 4-20 mm; inflorescences terminal leafy cymes
S. littoralis
31. Plants not having the above combination of characters
→ 32
32. Sepals 2-3(-4) mm; open flowers less than 5 mm diam.; petals not exceeding sepals, inconspicuous or absent
→ 33
32. Sepals 3.5 mm or longer; open flowers 5 mm or more diam.; petals equaling or exceeding sepals, conspicuous
→ 34
33. Capsules broadly ovoid to globose, to 1.5 times as long as broad; styles less than 1 mm; sepals in open flowers less than 2.5 mm, veins obscure (rarely 1)
S. calycantha
33. Capsules ovoid, more than 1.5 times as long as broad; styles longer than 1 mm; sepals in open flowers more than 2.5 mm with 1-3 prominent veins
S. borealis
34. Sepals 4.5-6 mm; leaf blades ovate to broadly lanceolate, coriaceous, apex spinous; capsules equaling sepals
S. ruscifolia
34. Sepals 3.5-5 mm; leaf blades lanceolate to linear-lanceolate, not strongly coriaceous, apex acute to acuminate, not spinous; capsules 1.5-2 times as long as sepals
S. longipes
Source FNA vol. 5, p. 110. FNA vol. 5, p. 96. Author: John K. Morton.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Stellaria Caryophyllaceae > subfam. Alsinoideae
Sibling taxa
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. longipes, S. media, S. neglecta, S. nitens, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
Subordinate taxa
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. longipes, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
Synonyms Alsine obtusa, Alsine viridula, Alsine washingtoniana, S. viridula, S. washingtoniana Alsine
Name authority Engelmann: Bot. Gaz. 7: 5. (1882) Linnaeus: Sp. Pl. 1: 421. (1753): Gen. Pl. ed. 5, 193. (1754)
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