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Goldie's starwort, long-stalk starwort

circumpolar starwort, prickly-leaf starwort

Habit Plants perennial, forming small to large clumps or mats, or diffuse, from slender rhizomes. Plants perennial, forming small to moderate clumps, from elongate rhizomes.
Stems

erect to straggling, branched or not, 4-angled, 3–32 cm, glabrous or softly pubescent, angles not minutely papillate-scabrid.

erect, branched, 4-angled, 3–20 cm, glabrous.

Leaves

sessile;

blade green, frequently glaucous, 1–3-veined, midrib prominent, lanceolate to linear-lanceolate, widest at base, 0.4–2.6(–4) cm × 1–4 mm, strongly coriaceous or not, base round, margins entire, convex, glabrous or ciliate, apex acute to acuminate, not spinescent, shiny, smooth, glabrous or sparingly villous, base usually glabrous, rarely with few cilia.

sessile;

blade ovate to broadly lanceolate, widest below middle, 0.4–2 cm × 2–6 mm, coriaceous, base round, margins entire, apex acuminate, spinous, glabrous.

Inflorescences

with flowers solitary, or terminal, 3–30-flowered (rarely more) cymes;

bracts lanceolate, 2–10 mm, herbaceous with scarious margins, or scarious throughout, glabrous or ciliate.

with flowers solitary, subterminal in axils of foliage leaves, or terminal, 2–5-flowered cymes;

bracts (when present) lanceolate, distally reduced, 3–14 mm, herbaceous, margins scarious, apex acuminate.

Pedicels

ascending to erect, straight, 5–30 mm, glabrous or softly pubescent.

stiffly erect, 5–40 mm, glabrous.

Flowers

5–10 mm diam.;

sepals 5, 3-veined, midrib prominent, lanceolate to ovate-lanceolate, 3.5–5 mm, margins convex, narrow, scarious, sometimes ciliate, apex acute, glabrous or pubescent;

petals 5, 3–8 mm, 1–1.5 times as long as sepals;

stamens 5–10;

styles 3(–6), ascending, curled at tip, ca. 1.5 mm.

10–13 mm diam.;

sepals 5, 3-veined, lanceolate, 4.5–6 mm, margins narrow, scarious, apex acute, glabrous or sparsely pilose;

petals 5, 5–7 mm, 1–1.5 times as long as sepals, blade apex with lobes oblanceolate;

stamens 10, in 2 whorls;

styles 3(–4), ascending and outwardly curved, 2 mm.

Capsules

blackish purple or straw colored, ovoid to ovoid-lanceoloid, 4–6 mm, 1.5–2 times as long as sepals, apex broadly acute, opening by 6 valves;

carpophore absent.

olive green, ovoid, 4–6 mm, equaling and enclosed in sepals, opening by 6(–8) valves;

carpophore absent.

Seeds

brown, reniform to globose, 0.6–0.9 mm diam., shallowly tuberculate to smooth.

brown, reniform-rotund, 0.8–1.2 mm, rugose.

2n

= 52–104, (107).

Stellaria longipes

Stellaria ruscifolia

Phenology Flowering summer.
Habitat Tundra, gravelly places
Elevation 0-1100 m (0-3600 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MI; MN; MT; ND; NM; NY; OR; SD; UT; WA; WY; AB; BC; MB; NB; NT; NU; ON; QC; SK; YT; Circumpolar
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from FNA
AK; Asia (Russian Far East)
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Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

North American material of Stellaria ruscifolia is variable but tends to be more compact and smaller than that from the Russian Far East. It is referable to subsp. aleutica. It appears to be a relative of S. longipes, and some forms of the latter with wider leaves (S. crassipes) are very similar but do not have the coriaceous, more or less prickly leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Capsules purplish black; stems variable, commonly compact, erect; leaf blades very variable, from linear-lanceolate to ovate-triangular
subsp. longipes
1. Capsules straw colored; stems elongate, straggling; leaf blades narrowly lanceolate
subsp. arenicola
Source FNA vol. 5, p. 108. FNA vol. 5, p. 113.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Stellaria Caryophyllaceae > subfam. Alsinoideae > Stellaria
Sibling taxa
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. longipes, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. umbellata
Subordinate taxa
S. longipes subsp. arenicola, S. longipes subsp. longipes
Synonyms Alsine longipes S. ruscifolia subsp. aleutica
Name authority Goldie: Edinburgh Philos. J. 6: 327. (1822) Pallas ex Schlectendal: Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 7: 194. (1816)
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