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Goldie's starwort, long-stalk starwort

common or grass-leaf stitchwort or starwort, common starwort, common stitchwort, grass-leaf starwort, grass-leaf stitchwort, grass-like starwort, mouron des champs

Habit Plants perennial, forming small to large clumps or mats, or diffuse, from slender rhizomes. Plants perennial, coarse, rhizomatous; rhizomes slender, elongate.
Stems

erect to straggling, branched or not, 4-angled, 3–32 cm, glabrous or softly pubescent, angles not minutely papillate-scabrid.

decumbent or ascending, straggling, diffusely branched, smoothly 4-angled, 20–90 cm, brittle, glabrous.

Leaves

sessile;

blade green, frequently glaucous, 1–3-veined, midrib prominent, lanceolate to linear-lanceolate, widest at base, 0.4–2.6(–4) cm × 1–4 mm, strongly coriaceous or not, base round, margins entire, convex, glabrous or ciliate, apex acute to acuminate, not spinescent, shiny, smooth, glabrous or sparingly villous, base usually glabrous, rarely with few cilia.

sessile;

blade linear-lanceolate to narrowly lanceolate, widest near base, 1.5–4 cm × 1–6 mm, base round, margins smooth, apex acute, often ciliate near base, otherwise glabrous, not glaucous.

Inflorescences

with flowers solitary, or terminal, 3–30-flowered (rarely more) cymes;

bracts lanceolate, 2–10 mm, herbaceous with scarious margins, or scarious throughout, glabrous or ciliate.

terminal, 5–many-flowered, open, conspicuously branched cymes;

bracts narrowly lanceolate, 1–5 mm, wholly scarious, margins ciliate, apex acuminate.

Pedicels

ascending to erect, straight, 5–30 mm, glabrous or softly pubescent.

divaricate, 10–30 mm, glabrous.

Flowers

5–10 mm diam.;

sepals 5, 3-veined, midrib prominent, lanceolate to ovate-lanceolate, 3.5–5 mm, margins convex, narrow, scarious, sometimes ciliate, apex acute, glabrous or pubescent;

petals 5, 3–8 mm, 1–1.5 times as long as sepals;

stamens 5–10;

styles 3(–6), ascending, curled at tip, ca. 1.5 mm.

5–12 mm diam., rarely larger;

sepals 5, distinctly 3-veined, narrowly lanceolate to triangular, 3–7 mm, margins narrow, straight, scarious, apex acute, glabrous;

petals 5, 3–7 mm, equaling or longer than sepals;

stamens 10, all, some, or none fully developed and fertile;

styles 3, ascending, ca. 3 mm.

Capsules

blackish purple or straw colored, ovoid to ovoid-lanceoloid, 4–6 mm, 1.5–2 times as long as sepals, apex broadly acute, opening by 6 valves;

carpophore absent.

green or straw colored, narrowly ovoid, 5–7 mm, longer than sepals, apex acute, opening by 3 valves, splitting into 6;

carpophore absent.

Seeds

brown, reniform to globose, 0.6–0.9 mm diam., shallowly tuberculate to smooth.

reddish brown, reniform-rotund, ca. 1 mm diam., rugose in concentric rings.

2n

= 52–104, (107).

= 39, 52.

Stellaria longipes

Stellaria graminea

Phenology Flowering late spring–early summer.
Habitat Rough grasslands, pastures, hayfields, roadsides
Elevation 0-1200 m (0-3900 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MI; MN; MT; ND; NM; NY; OR; SD; UT; WA; WY; AB; BC; MB; NB; NT; NU; ON; QC; SK; YT; Circumpolar
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[BONAP county map]
from FNA
CA; CO; CT; DC; IA; ID; IL; IN; KS; KY; MA; ME; MI; MN; MO; MT; NC; NH; NJ; NV; NY; OH; OR; PA; RI; SC; TN; VA; VT; WA; WI; WV; BC; NB; NL; NS; ON; PE; QC; SPM; Europe [Introduced in North America]
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Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Europe, both diploid and tetraploid cytotypes of Stellaria graminea occur with occasional triploid hybrids. Only the tetraploid form has been found in North America, except for a triploid colony in Newfoundland. This species is often confused with S. longifolia but differs in its stems, which are very angular, glabrous, and not scabrid; the narrowly triangular leaves on the flowering stems; the smooth leaf margins; the stiff, triangular, prominently 3-veined sepals; and the larger, rugulose seeds.

The sterile overwintering shoots of Stellaria graminea have broader elliptic to elliptic-lanceolate leaf blades measuring 5–15 × 1.5–4 mm. They are broadest near the middle. This state of the plant has been named var. latifolia Petermann. Usually S. graminea has perfect flowers but occasionally plants that are entirely staminate-sterile are encountered. The flowers in these are partially fertile depending on the occurrence of cross- pollination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Capsules purplish black; stems variable, commonly compact, erect; leaf blades very variable, from linear-lanceolate to ovate-triangular
subsp. longipes
1. Capsules straw colored; stems elongate, straggling; leaf blades narrowly lanceolate
subsp. arenicola
Source FNA vol. 5, p. 108. FNA vol. 5, p. 105.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Stellaria Caryophyllaceae > subfam. Alsinoideae > Stellaria
Sibling taxa
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. longipes, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
Subordinate taxa
S. longipes subsp. arenicola, S. longipes subsp. longipes
Synonyms Alsine longipes Alsine graminea
Name authority Goldie: Edinburgh Philos. J. 6: 327. (1822) Linnaeus: Sp. Pl. 1: 422. (1753)
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