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Goldie's starwort, long-stalk starwort

Habit Plants perennial, forming small to large clumps or mats, or diffuse, from slender rhizomes. Herbs, winter annual, annual, biennial, or perennial; taprooted and/or rhizomatous, rarely with tuberous thickenings (Pseudostellaria).
Stems

erect to straggling, branched or not, 4-angled, 3–32 cm, glabrous or softly pubescent, angles not minutely papillate-scabrid.

prostrate to ascending or erect, simple or branched.

Leaves

sessile;

blade green, frequently glaucous, 1–3-veined, midrib prominent, lanceolate to linear-lanceolate, widest at base, 0.4–2.6(–4) cm × 1–4 mm, strongly coriaceous or not, base round, margins entire, convex, glabrous or ciliate, apex acute to acuminate, not spinescent, shiny, smooth, glabrous or sparingly villous, base usually glabrous, rarely with few cilia.

opposite, connate proximally or not, often petiolate (basal leaves), not stipulate;

blade subulate or linear to spatulate, lanceolate, or broadly ovate, seldom succulent.

Inflorescences

with flowers solitary, or terminal, 3–30-flowered (rarely more) cymes;

bracts lanceolate, 2–10 mm, herbaceous with scarious margins, or scarious throughout, glabrous or ciliate.

terminal or axillary cymes, or flowers solitary;

bracts foliaceous or reduced, herbaceous to scarious (or rarely absent);

involucel bracteoles absent.

Pedicels

ascending to erect, straight, 5–30 mm, glabrous or softly pubescent.

present or rarely flowers sessile.

Flowers

5–10 mm diam.;

sepals 5, 3-veined, midrib prominent, lanceolate to ovate-lanceolate, 3.5–5 mm, margins convex, narrow, scarious, sometimes ciliate, apex acute, glabrous or pubescent;

petals 5, 3–8 mm, 1–1.5 times as long as sepals;

stamens 5–10;

styles 3(–6), ascending, curled at tip, ca. 1.5 mm.

bisexual or seldom unisexual, sometimes inconspicuous;

perianth and androecium hypogynous or perigynous, often slightly;

hypanthium cup-, dish-, or disc-shaped;

sepals (4–)5, distinct or seldom connate basally, sometimes hooded, not awned;

petals absent or (1–)4–5, usually white, sometimes translucent, yellowish white, pink, or brownish, seldom clawed, auricles absent, coronal appendages absent, blade apex entire or 2-fid, sometimes jagged or emarginate, rarely laciniate;

stamens absent or (1–)5(–10), in 1 or 2 whorls, arising from base of ovary, a nectariferous disc, or sometimes the hypanthium or hypanthium rim;

staminodes absent or 1–5(–8);

ovary 1- or rarely 3-locular (Wilhelmsia);

styles (2–)3–5(–6), distinct;

stigmas (2–)3–5(–6).

Fruits

capsules, or rarely utricles (Scleranthus), opening by (2–)3–6, occasionally 8 or 10 valves or (3 or) 6–10 teeth;

carpophore present or often absent.

Capsules

blackish purple or straw colored, ovoid to ovoid-lanceoloid, 4–6 mm, 1.5–2 times as long as sepals, apex broadly acute, opening by 6 valves;

carpophore absent.

Seeds

brown, reniform to globose, 0.6–0.9 mm diam., shallowly tuberculate to smooth.

1–60+, yellowish or tan to dark red or often brown or black, usually reniform or triangular to circular and laterally compressed or ovoid to globose, rarely oblong and dorsiventrally compressed (Holosteum);

embryo usually peripheral and curved, rarely central and straight (Holosteum).

x

= 6–15, 17–19, 23.

2n

= 52–104, (107).

Stellaria longipes

Caryophyllaceae subfam. alsinoideae

Distribution
from FNA
AK; AZ; CA; CO; ID; MI; MN; MT; ND; NM; NY; OR; SD; UT; WA; WY; AB; BC; MB; NB; NT; NU; ON; QC; SK; YT; Circumpolar
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[BONAP county map]
North-temperate regions; South America (Andean region); Europe (Mediterranean region); w Asia; c Asia (Himalayas, Mediterranean region); Africa (Mediterranean region)
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 30, species ca. 1040 (16 genera, 137 species in the flora).

Alsinoideae, often considered basal in the family and the least specialized, is in some ways the most heterogeneous of the subfamilies. Members of its largest tribe (Alsineae) share the following characteristics: stipules absent, sepals free or at most basally connate, and capsular fruits. Indehiscent fruits, relatively short hypanthia, and other floral reductions occur in varying combinations in the approximately 30 species placed in four other tribes. A broad molecular survey of Alsinoideae has revealed two major lineages and lack of support for the existing tribal circumscriptions (M. Nepokroeff et al. 2002). About three-fourths of the species are members of Arenaria, Cerastium, Minuartia, and Stellaria.

Attempts have been made to move Scleranthus (fruit a utricle surrounded by an enlarged hypanthium) from Alsinoideae to either Paronychioideae (J. Hutchinson 1973, as Illecebraceae) or Scleranthaceae (A. Takhtajan 1997). Recent molecular and morphological studies by R. D. Smissen et. al. (2002, 2003) supported its retention in the Alsinoideae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Capsules purplish black; stems variable, commonly compact, erect; leaf blades very variable, from linear-lanceolate to ovate-triangular
subsp. longipes
1. Capsules straw colored; stems elongate, straggling; leaf blades narrowly lanceolate
subsp. arenicola
Source FNA vol. 5, p. 108. FNA vol. 5, p. 50. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Stellaria Caryophyllaceae
Sibling taxa
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
Subordinate taxa
S. longipes subsp. arenicola, S. longipes subsp. longipes
Synonyms Alsine longipes
Name authority Goldie: Edinburgh Philos. J. 6: 327. (1822) Fenzl: in S. L. Endlicher, Gen. Pl. 13: 963. (1840)
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