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crisp sandwort, crisp starwort, crisped starwort, curled starwort, ruffled starwort

circumpolar starwort, prickly-leaf starwort

Habit Plants perennial, forming small to large mats, from slender rhizomes. Plants perennial, forming small to moderate clumps, from elongate rhizomes.
Stems

trailing to ascending, branched, 4-angled, 10–60 cm, glabrous.

erect, branched, 4-angled, 3–20 cm, glabrous.

Leaves

subsessile;

blade broadly elliptic to ovate, 0.4–2.6 cm × 2–15 mm, base round to cuneate, margins entire, apex acuminate, glabrous or with a few scattered cilia.

sessile;

blade ovate to broadly lanceolate, widest below middle, 0.4–2 cm × 2–6 mm, coriaceous, base round, margins entire, apex acuminate, spinous, glabrous.

Inflorescences

with flowers solitary in leaf axils;

bracts absent.

with flowers solitary, subterminal in axils of foliage leaves, or terminal, 2–5-flowered cymes;

bracts (when present) lanceolate, distally reduced, 3–14 mm, herbaceous, margins scarious, apex acuminate.

Pedicels

ascending, straight, mostly 5–30 mm, glabrous.

stiffly erect, 5–40 mm, glabrous.

Flowers

4–5 mm;

sepals 5, prominently 3-veined, lanceolate, 2–4 mm, margins broadly scarious, apex acute to acuminate, glabrous;

petals usually absent, rarely 1–5 and much shorter than sepals;

stamens 10 or fewer;

styles 3, spreading to ascending, curved but not curled, ca. 1 mm.

10–13 mm diam.;

sepals 5, 3-veined, lanceolate, 4.5–6 mm, margins narrow, scarious, apex acute, glabrous or sparsely pilose;

petals 5, 5–7 mm, 1–1.5 times as long as sepals, blade apex with lobes oblanceolate;

stamens 10, in 2 whorls;

styles 3(–4), ascending and outwardly curved, 2 mm.

Capsules

straw colored or brownish, ovoid to ovoid-ellipsoid, 3.5–6 mm, equaling or slightly exceeding sepals, apex broadly acute, opening by 6 valves;

carpophore absent.

olive green, ovoid, 4–6 mm, equaling and enclosed in sepals, opening by 6(–8) valves;

carpophore absent.

Seeds

brown, broadly elliptic, 0.7–1 mm (longest axis), distinctly rugose.

brown, reniform-rotund, 0.8–1.2 mm, rugose.

2n

= 26, 52.

Stellaria crispa

Stellaria ruscifolia

Phenology Flowering summer. Flowering summer.
Habitat Wet soil in woods, shaded streambanks and shores Tundra, gravelly places
Elevation 0-2300 m (0-7500 ft) 0-1100 m (0-3600 ft)
Distribution
from FNA
AK; CA; ID; MT; OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; Asia (Russian Far East)
[BONAP county map]
Discussion

North American material of Stellaria ruscifolia is variable but tends to be more compact and smaller than that from the Russian Far East. It is referable to subsp. aleutica. It appears to be a relative of S. longipes, and some forms of the latter with wider leaves (S. crassipes) are very similar but do not have the coriaceous, more or less prickly leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 104. FNA vol. 5, p. 113.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Stellaria Caryophyllaceae > subfam. Alsinoideae > Stellaria
Sibling taxa
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. longipes, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. ruscifolia, S. umbellata
S. alaskana, S. alsine, S. americana, S. borealis, S. calycantha, S. corei, S. crassifolia, S. crispa, S. cuspidata, S. dicranoides, S. fontinalis, S. graminea, S. holostea, S. humifusa, S. irrigua, S. littoralis, S. longifolia, S. longipes, S. media, S. neglecta, S. nitens, S. obtusa, S. pallida, S. palustris, S. parva, S. porsildii, S. pubera, S. umbellata
Synonyms Alsine crispa, S. borealis var. crispa S. ruscifolia subsp. aleutica
Name authority Chamisso & Schlechtendal: Linnaea 1: 51. (1826) Pallas ex Schlectendal: Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 7: 194. (1816)
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