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cow-horn peat-moss, sphagnum

rough peat moss, shaggy peat, sphagnum, spiky bog-moss, spread-leaf peat moss, spreading-leaf bog moss

Habit Plants small, slender, often wiry; green, yellow-brown or golden brown; capitulum small with terminal bud absent. Plants robust, stiff; green, pale green, yellow-green; large terminal bud; typically as loose carpets in coniferous forests.
Stem(s)

leaves triangular-lingulate, 0.8 mm or less;

apex entire or weakly denticulate, hyaline cells sometimes septate, efibrillose and aporose except near apex.

green to red-brown; 2–3 superficial cortical layers.;

stem leaves shorter than branch leaves, ovate-lingulate to oblong-lingulate, 1.6–1.8 × 1–1.2 mm;

hyaline cells mostly nonseptate.

Branches

often short and blunt.

long and tapering with distinct squarrose spreading leaves, often terete in tundra forms.

Branch leaves

larger than stem leaves, 1.9–2.8 mm, conspicuously squarrose from ovate-hastate base and abruptly narrowed 1/2–1/3 distance from apex into involute-concave acumen, often terete in tundra forms;

hyaline cells convex on both surfaces, non-ringed pores at ends and corners of cells, ringed pores on concave surface (4–8/cell) and nonringed pores (2–4/cell) on convex surface, internal commissural walls smooth or indistinctly papillose, chlorophyllous cells ovate triangular with widest part at or close to the convex surface.

Sexual condition

dioicous.

monoicous.

Capsule

exserted, with few pseudostomata.

Spores

30–35 µm; finely papillose on both surfaces, distinct bifurcated Y-mark sculpture on distal surface;

proximal laesurae more than 0.5 spore radius.

17–30 µm;

proximal surface finely papillose, distal surface smooth with raised bifurcated Y-mark sculpture;

proximal laesura more than 0.5 spore radius.

Branch

fascicles with 2–3 spreading and 2–3 pendent branches Branch leaves broadly ovate, subsecund, hyaline cells on convex surface with very numerous small pores (18–40 per cell) in a continuous row along the commissures, concave surface usually aporose.

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems with 1–2 layers of cortical cells.

Sphagnum subsecundum

Sphagnum squarrosum

Phenology Sporophytes abundant, capsules mature early to mid summer.
Habitat Minerotrophic, near the edges of open, poor fens, less commonly found in open medium fens Forming loose carpets in rich habitats such as wet coniferous forests, Thuja swamps, karrs, medium fens, and stream margins
Elevation low to high elevations low to high elevations
Distribution
from FNA
AK; CA; CO; CT; ID; IL; IN; MA; ME; MI; MN; MT; NH; NJ; NY; OH; OR; PA; RI; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; QC; YT; Greenland; Eurasia
[WildflowerSearch map]
from FNA
AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; ME; MI; MN; MT; NC; NH; NJ; NY; OR; PA; SD; TN; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia; Pacific Islands (New Zealand)
[WildflowerSearch map]
Discussion

Sporophytes are rare in Sphagnum subsecundum. This species is often associated with S. angustifolium, S. centrale, S. fimbriatum, S. flexuosum, S. palustre, and S. teres. The most widespread and common species of sect. Subsecunda, it exhibits considerable phenotypic plasticity in size. However, the stem leaves are always quite small in comparison to those of similar species. See also discussions under 52. S. contortum and 55. S. inexspectatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In its typical robust form with strongly squarrose branch leaves, Sphagnum squarrosum is unmistakeable. Smaller forms such as occur in the higher mountains may be difficult to identify accurately without careful examination of microscopic details. In the tundra there sometimes occur large, terete forms of S. squarrosum but these are usually considerably more robust than S. teres. See also discussion under 14. S. strictum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 84. FNA vol. 27, p. 59.
Parent taxa Sphagnaceae > Sphagnum > sect. Subsecunda Sphagnaceae > Sphagnum > sect. Squarrosa
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. crispum S. squarrosum var. imbricatum
Name authority Nees: in J. Sturm et al., Deutschl. Fl. 2(17): species 3. (1819) Crome: Samml. Deut. Laubm., 24. (1803)
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