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rough peat moss, shaggy peat, sphagnum, spiky bog-moss, spread-leaf peat moss, spreading-leaf bog moss

Habit Plants robust, stiff; green, pale green, yellow-green; large terminal bud; typically as loose carpets in coniferous forests. Plants fairly slender to moderate-sized, green; forming low dense hummocks.
Stem(s)

green to red-brown; 2–3 superficial cortical layers.;

stem leaves shorter than branch leaves, ovate-lingulate to oblong-lingulate, 1.6–1.8 × 1–1.2 mm;

hyaline cells mostly nonseptate.

leaves generally longer than branch leaves, 1.1–1.7 mm, lingulate to lingulate-spathulate, hyaline cells mostly non-septate.

Branches

long and tapering with distinct squarrose spreading leaves, often terete in tundra forms.

terete.

Branch leaves

larger than stem leaves, 1.9–2.8 mm, conspicuously squarrose from ovate-hastate base and abruptly narrowed 1/2–1/3 distance from apex into involute-concave acumen, often terete in tundra forms;

hyaline cells convex on both surfaces, non-ringed pores at ends and corners of cells, ringed pores on concave surface (4–8/cell) and nonringed pores (2–4/cell) on convex surface, internal commissural walls smooth or indistinctly papillose, chlorophyllous cells ovate triangular with widest part at or close to the convex surface.

1–1.4 mm, broadly ovate, with a narrow involute tip;

hyaline cells only slightly bulging on either surface, in proximal half of leaf aporose on convex surface and with large faint pores on concave surface;

internal commissural walls distinctly papillose; chlorophyllous cells elliptical to elliptical-triangular in transverse section, enclosed on both surfaces with the widest part in the leaf middle.

Sexual condition

monoicous.

dioicous.

Spores

17–30 µm;

proximal surface finely papillose, distal surface smooth with raised bifurcated Y-mark sculpture;

proximal laesura more than 0.5 spore radius.

ca. 31 µm, ornamented by small somewhat amalgamated granulae.

Branch

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems with 1–2 layers of cortical cells.

fascicles of 2 spreading and 1–2 hanging branches.;

branch stems with 1–2 layers of cortical cells.

Sphagnum squarrosum

Sphagnum mirum

Phenology Sporophytes abundant, capsules mature early to mid summer. Sporophytes abundant, capsules mature August.
Habitat Forming loose carpets in rich habitats such as wet coniferous forests, Thuja swamps, karrs, medium fens, and stream margins Ecology poorly known but probably quite minerotrophic
Elevation low to high elevations low elevations
Distribution
from FNA
AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; ME; MI; MN; MT; NC; NH; NJ; NY; OR; PA; SD; TN; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia; Pacific Islands (New Zealand)
[WildflowerSearch map]
from FNA
AK
Discussion

In its typical robust form with strongly squarrose branch leaves, Sphagnum squarrosum is unmistakeable. Smaller forms such as occur in the higher mountains may be difficult to identify accurately without careful examination of microscopic details. In the tundra there sometimes occur large, terete forms of S. squarrosum but these are usually considerably more robust than S. teres. See also discussion under 14. S. strictum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphagnum mirum has only been recently discovered and so far is known only from its type locality, where it was growing in a fen mixed with S. teres.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 59. FNA vol. 27, p. 58.
Parent taxa Sphagnaceae > Sphagnum > sect. Squarrosa Sphagnaceae > Sphagnum > sect. Squarrosa
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. squarrosum var. imbricatum
Name authority Crome: Samml. Deut. Laubm., 24. (1803) Flatberg & Thingsgaard: Bryologist 106: 501. (2003)
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