Sphagnum sect. Subsecunda |
Sphagnum lescurii |
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Lescur's sphagnum |
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Habit | Plants erect to prostrate, extremely variable, capitulum rarely well developed; green, yellowish, light brown, golden brown, reddish brown to dark brown. | Plants moderate-sized to robust; upright, prostrate, or aquatic; green, pale yellow, golden brown, dark brown, tinged with red in exposed sites and purplish in aquatic forms; capitulum rounded and often strongly twisted. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Stem(s) | green to dark brown, superficial cortex of 0–3 layers of efibrillose, non-ornamented, enlarged, thin-walled cells; cells in outer layer aporose or with single round to elliptical wall thinning adjacent to the distal cell wall, visible only with heavy staining.; stem leaves varying from smaller than to larger than branch leaves; triangular, ovate to lingulate; with rounded and sometimes erose apex; border entire; hyaline cells rhomboid to S-shaped, non-ornamented, efibrillose to fibrillose, aporose to sometimes porose, non- to multiply septate; neither surface resorbed. |
leaves lingulate to ovate-lingulate, 1.3–2 mm; apex truncate to rounded, usually denticulate; hyaline cells typically fibrillose for 1/2 of leaf or more, often 1–2-septate, convex surface with 4–12 or more pores per cell along the commissures, concave surface with fewer pores. |
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Branches | not always clearly dimorphic, spreading and pendent branches very similar. |
usually curving, often large and tumid. |
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Branch leaves | oval, ovate or ovate-lanceolate; hyaline cells fibrillose, non-ornamented; convex surface mostly with numerous elliptical to round pores (8–24 per cell) in rows along commissures on convex surface, concave surface with fewer or no pores; chlorophyllous cells elliptical in transverse section, ± equally exposed on both surfaces or slightly more on convex surface, end walls not thickened. |
broadly ovate to ovate-lanceolate, 1.3–2.5 mm, greatly elongated in aquatic forms, straight or infrequently subsecund or subsquarrose; hyaline cells with 10–22 pores along the commissures on the convex surface, no or fewer pores per cell (1–8) on the concave surface. |
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Sexual condition | dioicous. |
dioicous. |
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Capsule | with few pseudostomata. |
exserted, with few pseudostomata. |
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Spores | 22–41 µm, with or without raised surface sculpture on distal surface; proximal laesura more than 0.5 spore radius. |
27–34 µm; finely papillose on both surfaces, with distinct raised Y-mark sculpture (indistinctly bifurcated Y-mark) on the distal surface; proximal laesura less than 0.5 spore radius. |
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Branch | fascicles 1–3 spreading and 0–2(–4) pendent.; branch stems green, surrounded by 1 layer of efibrillose, non-ornamented, thin-walled, inflated cells, with solitary short-necked retort cells or with conspicuously necked retort cells, interspersed with primarily aporose rectangular-shaped cells. |
fascicles with 2(rarely 3) spreading and 1–2(–3) pendent branches. |
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Sphagnum sect. Subsecunda |
Sphagnum lescurii |
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Habitat | Weakly minerotrophic in a broad range of wetlands, often of an aquatic or periodically dried character | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | low to moderate elevations | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution | Worldwide except Antarctica |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NF; NS; Europe
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Discussion | Species 99 (13 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sporophytes are uncommon in Sphagnum lescurii, which may be the most phenotypically variable of all the North American Sphagnum species, and quite probably deserves some taxonomic splitting. The tremendous phenotypic plasticity of this species, however, makes it quite difficult to sort out the genotypic component of variability, and thus most sphagnologists since Warnstorf have avoided the temptation of splitting and have instead treated this as one very variable species. This is the approach maintained in this treatment. We have also chosen not to use the earlier name S. denticulatum because its type is a phenotypic morphotype not clearly assignable to the current concept of either S. auriculatum or S. lescurii (K. I. Flatberg, pers. comm.). Some of the American material assignable to S. lescurii is quite likely the same as the European species S. auriculatum, but much of our material is certainly not the same. Until more definitive data are available, we have chosen to continue to use the name S. lescurii. The large stem leaf will generally distinguish this from similar species of sect. Subsecunda. See also discussion under 55. S. inundatum and 61. S. platyphyllum. The names Sphagnum alabamae Warnstorf, S. aquatile Warnstorf, S. obesum (Wilson) Warnstorf, S. rufescens (Nees & Hornschuch) Warnstorf, and S. turgidulum Warnstorf also have been applied to this taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 78. | FNA vol. 27, p. 81. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | S. unranked Subsecunda, S. unranked Cavifolia, S. unranked Comatosphagnum, S. unranked Cyclophylla, S. section Hemitheca | S. orlandense, S. plicatum, S. wieboldtii | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Lindberg) Schimper: Syn. Musc. Eur. ed. 2, 2: 843. (1876) | Sullivant: in A. Gray, Manual ed. 2, 611. (1856) | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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