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Habit Plants moderate-sized to robust, with distinct capitulum; green,yellowish brown.
Stem(s)

leaves ovate, ovate-lingulate to lingulate;

with broad, fringed apex;

little or no border along margins or base;

hyaline cells rhomboid, efibrillose, non-ornamented, aporose, usually nonseptate, with resorption gaps on exterior surface.

Branches

strongly dimorphic, pendent branches thinner but about same length as spreading branches.

Branch leaves

strongly squarrose or spreading, ovate, ovate-hastate or ovate-lanceolate, margins entire, apex involute and smooth;

hyaline cells fibrillose;

with large, round pores at cell ends and along commissures, sometimes with faint papillae on interior walls; chlorophyllous cells ovate-triangular, elliptical to ovate-elliptical in transverse section, more broadly exposed on convex surface, end walls not thickened.

Sexual condition

monoicous or dioicous.

Capsule

2 mm or more, with scattered pseudomata.

Spores

less than 30 µm, both surfaces smooth to finely papillose;

proximal laesura more than 0.5 spore radius.

Branch

stems green, with cortex surrounded by 1–2 layers of efibrillose, non-ornamented, enlarged, thin-walled cells, some cells apically porose with inconspicuous necks.

Sphagnum sect. Squarrosa

Distribution
North America; Eurasia; Pacific Islands (New Zealand)
Discussion

Species 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branch leaves markedly truncate; chlorophyll cells ovate-triangular in transverse section, with the widest part at or near the convex surface.
S. tundrae
1. Branch leaves not truncate; chlorophyll cells elliptical to elliptical-ovate with the broadest part typically some distance from the convex surface
→ 2
2. Branch leaves strongly squarrose (terete in tundra forms), large (1.9-3 mm); hyaline cells of branch leaves with ringed elliptic pores on concave surface and unringed pores on convex surface; stem leaves 1/2-2/3 as long as branch leaves (1.1-1.9 mm)
S. squarrosum
2. Branch leaves imbricate (squarrose in shade forms), moderate in size (1-1.5 mm); hyaline cells of branch leaves with unringed pores on concave and convex surfaces; stem leaves as long or longer than branch leaves (1-1.7 mm)
→ 3
3. Branch leaf hyaline cells near leaf base on convex surface mostly aporose and on concave surface with large, faint pores; 1-2 hanging branches.
S. mirum
3. Branch leaf hyaline cells near leaf base on convex surface with large, round pores and on concave surface mostly aporose; 2-3 hanging branches.
S. teres
Source FNA vol. 27, p. 58.
Parent taxa Sphagnaceae > Sphagnum
Subordinate taxa
S. mirum, S. squarrosum, S. teres, S. tundrae
Synonyms S. unranked Squarrosa
Name authority (Russow) Schimper: Syn. Musc. Eur. ed. 2, 835. (1876)
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