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five-rank peat-moss, sphagnum

steere's sphagnum

Habit Plants moderate-sized, typically stiff and compact, capitulum usually hemispherical; green, grayish white, pale yellow, purplish red, may have a slight metallic luster when dry. Plants moderate-sized to large, compact and stiff-stemmed with upswept branches; dark green and brown, dark chesnut brown, brown to blackish brown, often with bluish tinge when dry; forming dense low to moderately tall hummocks.
Stem(s)

leaves triangular to triangular-lingulate, 1–1.3 mm, apex acute to slightly obtuse, border broad at base (more than 0.25 width);

hyaline cells narrowly rhomboid, mostly 0–1-septate and mostly efibrillose.

leaves to 1.2 × 0.8 mm; rarely hemiisophyllous;

hyaline cells mostly nonseptate, comb-lamellae usually absent, but sometimes weak.

Branches

usually strongly 5-ranked.

short and blunt-tipped, leaves spreading.

Branch leaves

ovate to ovate-lanceolate, 1.1–1.5 mm, concave, straight, apex slightly involute;

hyaline cells on convex surface with numerous oval to elliptic pores along commissures grading from small pores near apex to large round pores at base, concave surface with large round pores in proximal portions of leaf.

ovate to ovate-elliptical, 1.6–2.2 × 1 mm;

hyaline cells on proximal half of convex surface with round to elliptic pores along the commissures;

comb-lamellae on hyaline cell wall where overlying chlorophyllous cells; chlorophyllous cells broadly triangular in transverse section and well-enclosed on convex surface, end wall not thickened.

Sexual condition

monoicous or dioicous.

dioicous.

Capsule

unknown.

Spores

19–27 µm, finely papillose on proximal surface, pusticulate on distal surface;

proximal laesura less than 0.4 spore radius.

unknown.

Branch

fascicles with mostly 3 spreading and 1–2 pendent branches.

fascicles with 2 spreading and 1 pendent branch.;

branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the end walls of cortical cells, cortical cell walls usually with large round pores.

Sphagnum quinquefarium

Sphagnum steerei

Phenology Capsules mature mid summer.
Habitat Weakly minerotrophic and hygrophytic, wet mineral bedrock, damp coniferous humus along coast and in montane regions Commonly in firm-bottomed poor to medium fen vegetation, subarctic to arctic regions
Elevation low to high elevations low to moderate elevations
Distribution
from FNA
AK; CT; GA; MA; MD; ME; MN; NC; NH; NY; PA; TN; VA; VT; WV; BC; MB; NB; NF; NS; ON; QC; Eurasia
from FNA
AK; MB; NT; NU; QC; YT; e Asia
Discussion

Sporophytes are common in Sphagnum quinquefarium. This species is usually associated with S. capillifolium, S. girgensohnii, and S. russowii. No other species of sect. Acutifolia has the combination of quinquefarious branch leaves and three spreading branches per fascicle. Sphagnum rubiginosum has three spreading branches but the branch leaves are quite unranked and its lingulate stem leaf is quite distinct from the triangular stem leaf of S. quinquefarium. See also discussion under 86. S. talbotianum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphagnum steerei is usually easily recognized in the field by its dense growth habit and dark brown color. Its ecology is somewhat unclear due to taxonomic confusion with S. imbricatum in the strict sense and S. austinii (R. E. Andrus 1987). The latter is the only species in the imbricatum complex that occurs where S. steerei does. Contrary to the view of H. A. Crum (1997), S. steerei is quite distinct from S. affine and S. austinii. There is no range overlap with either of those species and the macroscopic appearance is also quite different.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 96. FNA vol. 27, p. 54.
Parent taxa Sphagnaceae > Sphagnum > sect. Acutifolia Sphagnaceae > Sphagnum > sect. Sphagnum
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. acutifolium var. quinquefarium, S. schofieldii S. imbricatum var. arcticum
Name authority (Lindberg) Warnstorf: Hedwigia 25: 222. (1886) R. E. Andrus: Bryologist 90: 218. (1987)
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