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Pylaes' sphagnum

lustrous peat-moss, sphagnum

Habit Plants slender and delicate, aquatic or prostrate, with a conspicuous terminal bud; dark greenish to purplish brown in submerged plants to deep salmon-red in prostrate plants, capitulum quite indistinct but with distinct terminal bud. Plants moderate-sized, relatively robust, smaller in exposed sites, elongated in shaded sites, soft, capitulum ± large; pale green, green, yellowish, yellow-brown, brown, pinkish, purplish; with strong metallic sheen when dry.
Stem(s)

leaves broadly ovate, 1.5–2(2.5) mm; straight;

hyaline cells fibrillose and nearly aporose, with single small pores occasionally found in the distal cell ends on the concave surface.

leaves triangular to triangular-lingulate, 1.2–1.7 mm, apex acute to sometimes shortly cuspidate due to involute margins near apex, border narrow at base (less than 0.25 width);

hyaline cells mostly efibrillose and aporose, 0–1(–2)-septate.

Branches

lacking or short and slender.

long and tapering, unranked.

Branch leaves

when present similar to stem leaves but smaller, 0.8–1.2 mm, hyaline cells fibrillose and mostly aporose, 1–6 irregularly round-shaped membrane gaps in some cells near apex on convex surface.

1.3–2.7 mm, ovate to ovate-lanceolate, 1.3–2.7 mm, concave, straight, apex involute;

hyaline cells on convex surface with 2–6 large, narrowly elliptic ringed pores along the commissures, but usually absent from marginal regions of leaf, concave surface aporose or with 1–4 large round pores per cell especially near base.

Sexual condition

dioicous.

monoicous.

Capsule

usually immersed in perichaetiale leaves, but may be slightly emergent, pseudostomata absent from capsule surface.

Spores

29–41 µm, coarsely papillose on both surfaces, indistinct raised sculpture on distal surface;

proximal laesura less than 0.5 spore radius.

22–32 µm, finely to irregularly coarsely-papillose on proximal surface, irregularly coarsely papillose on distal surface;

proximal laesura less than 0.5 spore radius.

Branch

fascicles none or 1 spreading branch.

fascicles with 2 spreading and 1–2 pendent branches.

Sphagnum pylaesii

Sphagnum subnitens

Phenology Capsules mature early summer.
Habitat Weakly minerotrophic, wet rocks, poor fens Coastal oceanic mires that range from weakly to moderately minerotrophic
Elevation low to high elevations low to moderate elevations
Distribution
from FNA
MA; ME; NC; NH; NJ; NF; NS; QC; Greenland; NY ; Europe
[WildflowerSearch map]
from FNA
AK; CA; OR; WA; BC; Eurasia; Pacific Islands (New Zealand)
Discussion

Sporophytes rare in Sphagnum pylaesii. This species is distributed as a pioneer on wet rocks associated with S. tenellum and S. papillosum, or in poor fens with S. pulchrum, S. majus, and S. papillosum. See also discussion under 53. S. cyclophyllum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sporophytes are common in Sphagnum subnitens. The ecology of this species is unclear because of past taxonomic confusion with S. subfulvum. However, the two clearly differ in gametophyte and spore morphology (K. I. Flatberg 1985; Cao T. and D. H. Vitt 1986). The metallic sheen and strongly pointed stem leaves will distinguish this species from the somewhat similar S. capillifolium. See also discussion under 83. S. subfulvum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 84. FNA vol. 27, p. 98.
Parent taxa Sphagnaceae > Sphagnum > sect. Subsecunda Sphagnaceae > Sphagnum > sect. Acutifolia
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms Hemitheca pylaiei, S. pylaesii var. austinii, S. pylaesii var. prostratum, S. pylaesii var. ramosum, S. sedoides, S. sedoides var. austinii, S. sedoides var. prostratum
Name authority Bridel: Bryol. Univ. 1: 749. (1827) Russow & Warnstorf: Verh. Bot. Vereins Prov. Brandenburg 30: 115, plate 3, fig. 9, plate 4, figs. 22, 23. (1888)
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