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Pylaes' sphagnum

fringe bogmoss, fringe peat-moss, sphagnum

Habit Plants slender and delicate, aquatic or prostrate, with a conspicuous terminal bud; dark greenish to purplish brown in submerged plants to deep salmon-red in prostrate plants, capitulum quite indistinct but with distinct terminal bud. Plants typically small and slender, larger and compact in the Arctic, capitulum small to moderate-sized, often with a conspicuous terminal bud; green, yellowish brown to brown; without metallic lustre when dry.
Stem(s)

leaves broadly ovate, 1.5–2(2.5) mm; straight;

hyaline cells fibrillose and nearly aporose, with single small pores occasionally found in the distal cell ends on the concave surface.

leaves spatulate to broad-spatulate, 0.8–1.5(–2) mm, strongly lacerate across the broad apex and often part way down the margins, border scarcely to strongly broadened at base (0.25 width of base or less);

hyaline cells rhomboid, efibrillose and often 1–2-septate.

Branches

lacking or short and slender.

not 5-ranked, quite terete, long, and slender Branch fascicles with 1– 2 spreading and 1–2 pendent branches.

Branch leaves

when present similar to stem leaves but smaller, 0.8–1.2 mm, hyaline cells fibrillose and mostly aporose, 1–6 irregularly round-shaped membrane gaps in some cells near apex on convex surface.

ovate to ovate-lanceolate; 1.1–1.5(–2) mm, slightly concave, straight;

apex involute;

margins entire;

hyaline cells on convex surface with numerous pores along the commissures grading from small pores near leaf apex to large pores at base, concave surface with large round pores at leaf apex and along margins.

Sexual condition

dioicous.

often monoicous.

Capsule

usually immersed in perichaetiale leaves, but may be slightly emergent, pseudostomata absent from capsule surface.

Spores

29–41 µm, coarsely papillose on both surfaces, indistinct raised sculpture on distal surface;

proximal laesura less than 0.5 spore radius.

20–27 µm, finely papillose on both surfaces;

proximal laesura less than 0.5 spore radius.

Branch

fascicles none or 1 spreading branch.

Sphagnum pylaesii

Sphagnum fimbriatum

Habitat Weakly minerotrophic, wet rocks, poor fens
Elevation low to high elevations
Distribution
from FNA
MA; ME; NC; NH; NJ; NF; NS; QC; Greenland; NY ; Europe
[WildflowerSearch map]
from USDA
North America; South America; Eurasia; Pacific Islands (New Zealand)
[WildflowerSearch map]
Discussion

Sporophytes rare in Sphagnum pylaesii. This species is distributed as a pioneer on wet rocks associated with S. tenellum and S. papillosum, or in poor fens with S. pulchrum, S. majus, and S. papillosum. See also discussion under 53. S. cyclophyllum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants small and slender, capitulum small and with a conspicuous terminal bud; stem leaves fimbriate down the sides and weakly to moderately bordered at the base
subsp. fimbriatum
1. Plants moderate-sized and compact, capitulum moderate sized and lacking a conspicuous terminal bud; stem leaves entire down the sides and strongly bordered at the base
subsp. concinnum
Source FNA vol. 27, p. 84. FNA vol. 27, p. 92.
Parent taxa Sphagnaceae > Sphagnum > sect. Subsecunda Sphagnaceae > Sphagnum > sect. Acutifolia
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Subordinate taxa
S. fimbriatum subsp. concinnum, S. fimbriatum subsp. fimbriatum
Synonyms Hemitheca pylaiei, S. pylaesii var. austinii, S. pylaesii var. prostratum, S. pylaesii var. ramosum, S. sedoides, S. sedoides var. austinii, S. sedoides var. prostratum
Name authority Bridel: Bryol. Univ. 1: 749. (1827) Wilson & Hooker: in J. D. Hooker, Fl. Antarct., 398. (1847)
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