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Pylaes' sphagnum

imbricate bogmoss, sphagnum

Habit Plants slender and delicate, aquatic or prostrate, with a conspicuous terminal bud; dark greenish to purplish brown in submerged plants to deep salmon-red in prostrate plants, capitulum quite indistinct but with distinct terminal bud. Plants moderate-sized and lax to somewhat compact, ± stiff-stemmed; moderate-sized to large, forming lawns or low, loose hummocks; green, yellow-brown to golden brown and often tinged with brown to purplish brown; capitulum ± flat in lax open-grown forms to ± rounded and compact in open-grown forms.
Stem(s)

leaves broadly ovate, 1.5–2(2.5) mm; straight;

hyaline cells fibrillose and nearly aporose, with single small pores occasionally found in the distal cell ends on the concave surface.

leaves to 1.3–1.9 × 0.6–1.2 mm; rarely hemiisophyllous;

hyaline cells non-ornamented, nonseptate or sometimes septate.

Branches

lacking or short and slender.

± tapering, leaves loosely imbricate to spreading and often squarrose in shade forms.

Branch leaves

when present similar to stem leaves but smaller, 0.8–1.2 mm, hyaline cells fibrillose and mostly aporose, 1–6 irregularly round-shaped membrane gaps in some cells near apex on convex surface.

ovate to ovate elliptical, 1.5–2 × 0.9–1.6 mm;

hyaline cells on convex surface with elliptic to more often round pores along the commissures, comb-lamellae can be present, but often absent or restricted to leaf bases; chlorophyllous cells broadly triangular in transverse section and well-enclosed on the convex surface;

end wall not thickened.

Sexual condition

dioicous.

dioicous.

Capsule

usually immersed in perichaetiale leaves, but may be slightly emergent, pseudostomata absent from capsule surface.

with scattered pseudostomata.

Spores

29–41 µm, coarsely papillose on both surfaces, indistinct raised sculpture on distal surface;

proximal laesura less than 0.5 spore radius.

27–31 µm; granulate on both surfaces;

laesura on proximal surface less than 0.5 the spore radius.

Branch

fascicles none or 1 spreading branch.

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems with cortical cell comb-lamellae weakly differentiated on interior wall, no or weak funnel-like projections on the interior end walls, pores in superficial wall mostly restricted to leaf attachments.

Sphagnum pylaesii

Sphagnum affine

Phenology Capsules fairly common, mature early to late summer.
Habitat Weakly minerotrophic, wet rocks, poor fens Widespread and often ruderal, wide variety of minerotrophic wetlands, especially abundant in forested mires
Elevation low to high elevations low to moderate elevations
Distribution
from FNA
MA; ME; NC; NH; NJ; NF; NS; QC; Greenland; NY ; Europe
[WildflowerSearch map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IL; IN; KY; LA; MA; MD; ME; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WV; NF; NS; PE; Europe
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Discussion

Sporophytes rare in Sphagnum pylaesii. This species is distributed as a pioneer on wet rocks associated with S. tenellum and S. papillosum, or in poor fens with S. pulchrum, S. majus, and S. papillosum. See also discussion under 53. S. cyclophyllum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphagnum affine may occur elsewhere but the taxonomy is unclear (K. I. Flatberg 1984). Although species of sect. Sphagnum are notoriously difficult to tell apart in the field, S. affine is typically smaller than S. centrale, S. palustre, and S. papillosum, the other brown species of this section with which it may occur. It is also much more likely to have somewhat squarrose branch leaves, especially in shade forms.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 84. FNA vol. 27, p. 49.
Parent taxa Sphagnaceae > Sphagnum > sect. Subsecunda Sphagnaceae > Sphagnum > sect. Sphagnum
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms Hemitheca pylaiei, S. pylaesii var. austinii, S. pylaesii var. prostratum, S. pylaesii var. ramosum, S. sedoides, S. sedoides var. austinii, S. sedoides var. prostratum S. imbricatum subsp. affine, S. imbricatum var. affine, S. imbricatum var. laeve
Name authority Bridel: Bryol. Univ. 1: 749. (1827) Renauld & Cardot: Rev. Bryol. 12: 44. (1885)
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