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fat peat-moss, papillose sphagnum

wulf's sphagnum

Habit Plants moderate-sized to fairly robust; strong-stemmed and generally compact, capitulum usually not much enlarged; greenish brown to deep golden brown; forming compact carpets in floating mats and depressions as well as dense stands on hummock sides and low hummocks.
Stem(s)

leaves to 1.3 × 0.7 mm; rarely hemiisophyllous;

hyaline cells non-ornamented, mostly septate.

Branches

generally short and blunt, leaves spreading.

Branch leaves

broadly ovate, 1.7 × 1 mm;

hyaline cells on convex surface with round to elliptic pores along the commissures, hyaline cell walls covered with papillae where overlying chlorophyllous cells; chlorophyllous cells trapezoidal to truncate-elliptic in transverse section, equally exposed on both surfaces or less exposed on convex surface, end walls thickened.

with chlorophyllous cells often with faint papillae on interior walls.

Sexual condition

dioicous.

Capsule

with numerous pseudostomata.

Spores

26–36 µm; more roughly papillose on distal surface than proximal surface, distinct raised, bifurcated-Y mark sculpture on distal surface;

proximal laesura 0.5 spore radius or more.

Branch

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the interior end walls, mostly with 1 pore per cell on superficial cell wall.

Sphagnum papillosum

Sphagnum wulfianum

Phenology Capsules mature mid to late summer.
Habitat Very common in very poor to poor fen mire habitats where it is often a major peat former, but scarce to absent in truly ombrotrophic peatlands sites Coniferous forests, and occasionally in Alnus or Salix karrs
Elevation low to moderate elevations low to moderate elevations
Distribution
from FNA
AK; CA; CT; DE; IL; IN; MD; ME; MI; MN; NC; NH; NJ; NY; OH; OR; PA; SC; VT; WA; WI; WV; BC; NB; NF; NS; ON; PE; QC; YT; Eurasia
[WildflowerSearch map]
from FNA
CT; IL; IN; MA; ME; MI; MN; NH; NY; OH; PA; RI; VT; WI; AB; BC; MB; NB; NF; NS; ON; QC; Greenland; Eurasia
Discussion

Sphagnum papillosum is often easily field-identifiable by its rich golden brown to dark brown color and short, blunt branches. Nearly all specimens have the papillae on the branch leaf chorophyll cells but a few smooth forms have been found. Such forms will have stem leaves with divided hyaline cells whereas in the confusable species S. palustre and S. centrale such cells are rare or absent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The sporophytes of Sphagnum wulfianum are moderately common. This is the most dry-growing species in North America, typically growing in association with Sphagnum centrale, S. girgensohnii, S. russowii, and S. squarrosum. It is easily recognized as the only species that regularly has more than six branches per fascicle. The Lycopodium clavatum-like growth habit and conifer swamp habitat along with the strongly 5-ranked branch leaves make it even easier to recognize in the field.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 53. FNA vol. 27, p. 85.
Parent taxa Sphagnaceae > Sphagnum > sect. Sphagnum Sphagnaceae > Sphagnum > sect. Polyclada
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii
Synonyms S. cymbifolium var. papillosum, S. papillosum var. laeve, S. papillosum var. sublaeve, S. waghornei
Name authority Lindberg: Acta Soc. Sci. Fenn. 10: 280. (1872) Girgensohn: Arch. Naturk. Liv- Ehst- Kurlands, Ser. 2, Biol. Naturk. 2: 173. (1860)
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