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sphagnum

steere's sphagnum

Habit Plants moderate-sized, soft and lax when wet, stiff when dry, typically very compact, capitulum flat and usually large; pale whitish, yellowish or purplish, occasionally a deep purple-red; without metallic sheen when dry. Plants moderate-sized to large, compact and stiff-stemmed with upswept branches; dark green and brown, dark chesnut brown, brown to blackish brown, often with bluish tinge when dry; forming dense low to moderately tall hummocks.
Stem(s)

leaves quite variable in shape, elongate-lingulate to ovate, broadest above the middle, 1.9–2.5 mm, slightly concave, straight;

apex broad and toothed;

hyaline cells narrowly rhomboid, 0–1-septate, distal portion fibrillose, convex surface with membrane pleats, concave surface with 1(2–3) oblong membrane gaps.

leaves to 1.2 × 0.8 mm; rarely hemiisophyllous;

hyaline cells mostly nonseptate, comb-lamellae usually absent, but sometimes weak.

Branches

rarely 5-ranked.

short and blunt-tipped, leaves spreading.

Branch leaves

ovate, 1.6–2.2 mm, concave, straight;

apex stiffly involute and broadly truncate with up to 8 teeth, border denticulate due to cell wall resorption and projecting cell walls;

hyaline cells strongly bulging on convex surface and nearly plane on the concave surface, convex surface with narrowly elliptic pores along commissures grading from smaller pores near the apex to large rounded pores at base, concave surface with large round pores in proximal regions of leaf.

ovate to ovate-elliptical, 1.6–2.2 × 1 mm;

hyaline cells on proximal half of convex surface with round to elliptic pores along the commissures;

comb-lamellae on hyaline cell wall where overlying chlorophyllous cells; chlorophyllous cells broadly triangular in transverse section and well-enclosed on convex surface, end wall not thickened.

Sexual condition

monoicous.

dioicous.

Capsule

unknown.

Spores

27–33 µm, finely papillose on both surfaces with distinct bifurcated Y-mark sculpture on distal surface;

proximal laesura less than 0.5 spore radius.

unknown.

Branch

fascicles with 2 spreading and 1–2 pendent branches.

fascicles with 2 spreading and 1 pendent branch.;

branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the end walls of cortical cells, cortical cell walls usually with large round pores.

Sphagnum molle

Sphagnum steerei

Phenology Capsules mature early to mid summer.
Habitat Weakly minerotrophic and hygrophytic, poor fens and sand dunes, forming tight cushions among grasses and sedges in savannas, pine barrens, swamps, pond margins, and ditches where periodic dessication is common Commonly in firm-bottomed poor to medium fen vegetation, subarctic to arctic regions
Elevation low to high elevations low to moderate elevations
Distribution
from FNA
AL; DE; FL; GA; KY; LA; ME; MS; NC; NJ; NY; SC; TX; VA; LB; Europe
from FNA
AK; MB; NT; NU; QC; YT; e Asia
Discussion

The sporophytes of Sphagnum molle are common. This species is usually easily distinguished from other red species of sect. Acutifolia by its relatively large, straight, loosely spreading and unranked branch leaves. Sphagnum tenerum, the other red species of sect. Acutifolia to which it is most similar, has branch leaves that are quite imbricate. Microscopically, the denticulate-margined branch leaves are unmistakeable.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphagnum steerei is usually easily recognized in the field by its dense growth habit and dark brown color. Its ecology is somewhat unclear due to taxonomic confusion with S. imbricatum in the strict sense and S. austinii (R. E. Andrus 1987). The latter is the only species in the imbricatum complex that occurs where S. steerei does. Contrary to the view of H. A. Crum (1997), S. steerei is quite distinct from S. affine and S. austinii. There is no range overlap with either of those species and the macroscopic appearance is also quite different.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 95. FNA vol. 27, p. 54.
Parent taxa Sphagnaceae > Sphagnum > sect. Acutifolia Sphagnaceae > Sphagnum > sect. Sphagnum
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. labradorense, S. tabulare S. imbricatum var. arcticum
Name authority Sullivant: Musc. Allegh., 205. (1846) R. E. Andrus: Bryologist 90: 218. (1987)
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