Sphagnum molle |
Sphagnum russowii |
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sphagnum |
Russow's peat-moss, Russow's sphagnum |
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Habit | Plants moderate-sized, soft and lax when wet, stiff when dry, typically very compact, capitulum flat and usually large; pale whitish, yellowish or purplish, occasionally a deep purple-red; without metallic sheen when dry. | Plants ± moderate-sized, stiff and open, compact on exposed sites, capitulum flat-topped and often stellate; green or variegated red and green, lacking metallic sheen when dry. |
Stem(s) | leaves quite variable in shape, elongate-lingulate to ovate, broadest above the middle, 1.9–2.5 mm, slightly concave, straight; apex broad and toothed; hyaline cells narrowly rhomboid, 0–1-septate, distal portion fibrillose, convex surface with membrane pleats, concave surface with 1(2–3) oblong membrane gaps. |
leaves lingulate, 1.3–1.6 mm, apex broadly rounded or pointed and notched (sometimes denticulate), border strong and broadened at base (more than 0.25 width); hyaline cells short sinuoid-rhombic, mostly efibrillose, 0–1(–2)-septate. |
Branches | rarely 5-ranked. |
long and slender, never 5-ranked. |
Branch leaves | ovate, 1.6–2.2 mm, concave, straight; apex stiffly involute and broadly truncate with up to 8 teeth, border denticulate due to cell wall resorption and projecting cell walls; hyaline cells strongly bulging on convex surface and nearly plane on the concave surface, convex surface with narrowly elliptic pores along commissures grading from smaller pores near the apex to large rounded pores at base, concave surface with large round pores in proximal regions of leaf. |
ovate-lanceolate, 1.3–1.6 mm, concave, straight, apex strongly involute; hyaline cells on convex surface with numerous round to elliptic pores along the commissures, grading from small round pores near the apex to large elliptic pores at the base, concave surface usually with large round pores throughout, but sometimes restricted to proximal portions of leaf. |
Sexual condition | monoicous. |
dioicous, but some specimens apparently monoicous. |
Spores | 27–33 µm, finely papillose on both surfaces with distinct bifurcated Y-mark sculpture on distal surface; proximal laesura less than 0.5 spore radius. |
18–33 µm, coarsely papillose on both surfaces; proximal laesura more than 0.5 spore radius. |
Branch | fascicles with 2 spreading and 1–2 pendent branches. |
fascicles with 2 spreading and 1–2 pendent branches. |
Sphagnum molle |
Sphagnum russowii |
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Phenology | Capsules mature early to mid summer. | Capsules mature late spring to early summer. |
Habitat | Weakly minerotrophic and hygrophytic, poor fens and sand dunes, forming tight cushions among grasses and sedges in savannas, pine barrens, swamps, pond margins, and ditches where periodic dessication is common | Minerotrophic and shade-tolerant, common on the margins of mires, open portions of poor to rich fens, up through timberline in montane regions in wet coniferous forests |
Elevation | low to high elevations | low to high elevations |
Distribution |
AL; DE; FL; GA; KY; LA; ME; MS; NC; NJ; NY; SC; TX; VA; LB; Europe |
AK; CA; CO; CT; DE; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; NH; NJ; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia
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Discussion | The sporophytes of Sphagnum molle are common. This species is usually easily distinguished from other red species of sect. Acutifolia by its relatively large, straight, loosely spreading and unranked branch leaves. Sphagnum tenerum, the other red species of sect. Acutifolia to which it is most similar, has branch leaves that are quite imbricate. Microscopically, the denticulate-margined branch leaves are unmistakeable. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sporophytes are uncommon in Sphagnum russowii. This species is associated with S. centrale, S. fallax, S. fimbriatum, S. girgensohnii, and S. squarrosum. Because of its not particularly distinct phenotype as well its strong tendency to produce hemiisophyllous stem leaves, S. russowii is probably the most frequently misidentified Sphagnum species. The combination of the flat, stellate capitulum, unranked branch leaves, and lingulate stem leaf will usually suffice to identify it. Sphagnum capillifolium has a rounded capitulum and a pointed stem leaf while S. subtile also has a rounded capitulum but a shorter and more triangular stem leaf. In montane and arctic mires it can be confused with S. warnstorfii but the latter usually has conspicuously 5-ranked branch leaves. As one might expect in such a widespread and common species, the characters can vary considerably. For example, one regularly finds plants that are consistent in every respect with the description except that they lack stem cortical pores. Some stem leaves have almost no septations in the hyaline cells while other forms have most of the cells septate. As with similar variation in the likewise common S. fuscum, there is no consistent pattern and so taxonomic recognition of the variants is unwarranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 95. | FNA vol. 27, p. 97. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. labradorense, S. tabulare | S. acutifolium var. robustum |
Name authority | Sullivant: Musc. Allegh., 205. (1846) | Warnstorf: Hedwigia 25: 225. (1886) |
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