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Mendocino sphagnum

flat-leaf peat-moss, sphagnum

Habit Plants moderately robust and lax; terminal bud somewhat enlarged; yellow-green to light brownish green. Plants small to moderate-sized, unbranched or sparsely branched; green, golden brown to brown, capitulum small with a large and conspicuous terminal bud.
Stem(s)

leaves broadly oblong-triangular, 1.2–1.5 mm; mostly appressed to stem;

apex obtuse;

hyaline cells narrow, usually nonseptate, efibrillose and aporose on convex surface near apex, on concave surface usually efibrillose with irregular pores along commissures in distal portion of leaf.

leaves broadly ovate, 1.2–2.2 mm; straight;

apex rounded;

hyaline cells non-septate, convex surface with numerous small pores (less than or equal to 1/6 cell diameter) forming a continuous row along the commissures, concave surface aporose or with a few scattered pores along the commissures and cell ends.

Branches

with loosely imbricate leaves; often 5-ranked;

leaves little to somewhat elongated at distal end.

short and blunt, sometimes lacking completely and plants simplex.

Branch leaves

ovate, ovate-lanceolate to lanceolate; 2–3.5 mm;

flat and undulate at margins and recurved at apex when dry; straight;

margins entire;

hyaline cells on concave surface with very numerous, small ringed or unringed pores along the commissures, 5–12 in distal portion of leaf and 20–27 in proximal portion, convex surface with 5–15 pores per cell in distal portion of leaf and 14–21 in proximal portion, pores usually without a ring; chlorophyllous cells triangular to trapezoidal in transverse section and exposed slightly on concave surface.

broadly ovate, 1.4–2.5(–3) mm; straight;

apex rounded;

hyaline cells as in stem leaf.

Sexual condition

dioicous.

dioicous.

Capsule

exserted, with few pseudostomata.

Spores

ca. 30 µm;

very slightly roughened.

23–35 µm;

papillose on both surfaces, with indistinct Y-mark sculpture on distal surface;

proximal laesura 0.5 spore radius or less.

Branch

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems green, with cortical cells in 1 layer with conspicuous necks.

fascicles of 1–3 branches, 1–2 of these spreading, branches usually not numerous.

Sphagnum mendocinum

Sphagnum platyphyllum

Phenology Capsules mature late spring to early summer.
Habitat Submerged or floating in weakly minerotrophic wet depressions of alder swamps, coniferous swamps, Spiraea thickets, sedge fens, raised bogs, and drainage ditches in mires Typically growing in minerotrophic habitats such as shores of lakes, ponds, streams, flarks of string mires, margins of open fens, especially seasonally flooded sites
Elevation low to moderate elevations low to high elevations
Distribution
from FNA
AK; CA; ID; MT; OR; WA; BC
from FNA
AK; AZ; CA; CO; CT; IN; MA; ME; MI; MN; MT; NH; NJ; NY; OH; OR; RI; VT; WI; WY; AB; BC; MB; NB; NF; ON; QC; YT; South America; Greenland; Eurasia
Discussion

Sporophytes of Sphagnum mendocinum are uncommon. Although the unique branch leaf porosity makes it unmistakable microscopically, it may be confused with other species in the field. It is unusual ecologically in that it seems to overlap both the carpet- and lawn-forming habits—in other words it seems intermediate between S. recurvum and S. cuspidatum, in the broad sense. Among the species that it overlaps floristically, it is more robust and darker colored than S. pacificum. The latter also has an apiculate stem leaf compared to the more or less obtuse stem leaves of S. mendocinum. The more wet-growing S. majus var. majus and S. majus var. norvegicum, with which it slightly overlaps in habitat, have branch leaves that are strongly elongated at the distal branch ends whereas those of S. mendocinum are not. See also discussion under 40. S. obtusum. Sexual condition and spore characters are taken from from H. A. Crum (1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sporophytes are uncommon in Sphagnum platyphyllum, which can be quite variable in size and the development of branch fascicles, with some forms even being simplex while other forms may have up to three branches per fascicle. The species can usually be recognized by the large stem leaves that are spreading and easily visible because of the scarcity of hanging branches. It should also be noted that in this species and S. contortum the 2–3-layered stem cortex is not an entirely consistent trait, as plants quite typical in all other respects are occasionally found with the cortex only 1-layered, at least in part.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 72. FNA vol. 27, p. 83.
Parent taxa Sphagnaceae > Sphagnum > sect. Cuspidata Sphagnaceae > Sphagnum > sect. Subsecunda
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. mendocinum var. gracilescens, S. mendocinum var. recurvum, S. mendocinum var. robustum S. laricinum var. platyphyllum, S. contortum var. platyphyllum, S. subsecundum var. platyphyllum
Name authority Sullivant: Icon. Musc., suppl.: 12. (1874) (Lindberg) Warnstorf: Flora 67: 481. (1884)
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