Sphagnum mendocinum |
Sphagnum girgensohnii |
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Mendocino sphagnum |
common green peat-moss, Girgensohn's sphagnum |
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Habit | Plants moderately robust and lax; terminal bud somewhat enlarged; yellow-green to light brownish green. | Plants moderate-sized to robust, open, very stiff and slender, less frequently compact, capitulum large, flat, and stellate; typically deep green in shaded sites to yellowish brown in more open sites; without metallic lustre when dry. |
Stem(s) | leaves broadly oblong-triangular, 1.2–1.5 mm; mostly appressed to stem; apex obtuse; hyaline cells narrow, usually nonseptate, efibrillose and aporose on convex surface near apex, on concave surface usually efibrillose with irregular pores along commissures in distal portion of leaf. |
leaves lingulate, broadly lingulate to lingulate-spatulate; 0.8–1.3 mm, apex broad, truncate and lacerate, border broad at base (more than 0.25 of base); hyaline cells rhomboid, efibrillose, and rarely septate. |
Branches | with loosely imbricate leaves; often 5-ranked; leaves little to somewhat elongated at distal end. |
typically long and tapering, not 5-ranked. |
Branch leaves | ovate, ovate-lanceolate to lanceolate; 2–3.5 mm; flat and undulate at margins and recurved at apex when dry; straight; margins entire; hyaline cells on concave surface with very numerous, small ringed or unringed pores along the commissures, 5–12 in distal portion of leaf and 20–27 in proximal portion, convex surface with 5–15 pores per cell in distal portion of leaf and 14–21 in proximal portion, pores usually without a ring; chlorophyllous cells triangular to trapezoidal in transverse section and exposed slightly on concave surface. |
ovate to ovate-lanceolate, 1–1.4(–1.8) mm, concave, straight, apex strongly involute, margins entire; hyaline cells on convex surface with numerous elliptic pores along the commissures, grading from small pores near the apex to large pores near the base, concave surface with large round pores along the margins and base. |
Sexual condition | dioicous. |
dioicous. |
Spores | ca. 30 µm; very slightly roughened. |
21–27 µm, moderately to coarsely papillose on both surfaces; proximal laesura less than 0.5 spore radius. |
Branch | fascicles with 2 spreading and 2–3 pendent branches.; branch stems green, with cortical cells in 1 layer with conspicuous necks. |
fascicles with 2 spreading and 1–2 pendent branches.; branch stem with solitary retort cells or in groups of 2–3, necks moderately distinct. |
Sphagnum mendocinum |
Sphagnum girgensohnii |
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Phenology | Capsules mature late summer. | |
Habitat | Submerged or floating in weakly minerotrophic wet depressions of alder swamps, coniferous swamps, Spiraea thickets, sedge fens, raised bogs, and drainage ditches in mires | Shade tolerant, forming carpets on moist forest floors, along small streams, up through subalpine zone |
Elevation | low to moderate elevations | low to high elevations |
Distribution |
AK; CA; ID; MT; OR; WA; BC |
AK; CA; CO; CT; ID; IL; IN; MA; MD; ME; MI; MN; NC; ND; NH; NJ; NY; OH; OR; PA; RI; TN; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia
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Discussion | Sporophytes of Sphagnum mendocinum are uncommon. Although the unique branch leaf porosity makes it unmistakable microscopically, it may be confused with other species in the field. It is unusual ecologically in that it seems to overlap both the carpet- and lawn-forming habits—in other words it seems intermediate between S. recurvum and S. cuspidatum, in the broad sense. Among the species that it overlaps floristically, it is more robust and darker colored than S. pacificum. The latter also has an apiculate stem leaf compared to the more or less obtuse stem leaves of S. mendocinum. The more wet-growing S. majus var. majus and S. majus var. norvegicum, with which it slightly overlaps in habitat, have branch leaves that are strongly elongated at the distal branch ends whereas those of S. mendocinum are not. See also discussion under 40. S. obtusum. Sexual condition and spore characters are taken from from H. A. Crum (1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sporophytes are uncommon in Sphagnum girgensohnii. This species is most frequently associated with S. russowii, but also found growing with S. centrale, S. fallax, S. fimbriatum, S. warnstorfii, and S. magellanicum when growing in shaded sites of mires. It is very similar to S. rubiginosum, but S. girgensohnii lacks any reddish pigments, has only 2 spreading branches per fascicle, infrequently produces sporophytes, and differs in spore morphology. Throughout much of its range, S. girgensohnii is readily recognized by its green color and its large, slender, strongly stellate capitulum. In the more northern portion of its range, it frequently forms compact stands with a golden brown color and then the stem leaf must often be examined for accurate identification. In Alaska it overlaps morphogically with S. fimbriatum subsp. concinnum, which can look very similar but will have a more spatulate stem leaf that is lacerate completely across the broad flat apex and slightly down the sides. Sphagnum girgensohnii, on the other hand, has stem leaves only lacerate for about 3/4 of the apex width and less conspicuously broadened at the apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 72. | FNA vol. 27, p. 94. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. mendocinum var. gracilescens, S. mendocinum var. recurvum, S. mendocinum var. robustum | S. mehneri |
Name authority | Sullivant: Icon. Musc., suppl.: 12. (1874) | Russow: Beitr. Torfm., 46. (1865) |
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