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olive peat-moss, sphagnum

sphagnum

Habit Plants moderate-sized to robust, fairly weak-stemmed, lax in submersed forms, ± sprawling in emergent froms; golden brown to dark brown; capitulum weakly 5-radiate, branches straight to strongly laterally curved. Plants moderate-sized, pale green, yellow-green to occasionally strongly reddish; growing in loose mats.
Stem(s)

leaves triangular-lingulate, 0.8–1.4 mm, spreading to appressed;

apex acute to narrowly obtuse, hyaline cells nonseptate and fibrillose near apex.

leaves very small, less than 0.8 mm, triangular with blunt rounded apex.

Branches

unranked or weakly 5-ranked, straight to strongly curved, leaves moderately elongated at distal end.

erect in distal portion of plants.

Branch leaves

ovate-lanceolate to narrowly ovate-lanceolate, 1.8–3.4 mm; straight to strongly subsecund; weakly undulate and recurved when dry;

margins entire;

hyaline cells on convex surface with 1–2 free pores per fibril interval, concave surface aporose or rarely with a few wall thinnings in cell ends and angles; chlorophyllous cells trapezoidal in transverse section and narrowly exposed on concave surface.

large, 2.8 mm or longer, sub-squarrose, ovate, involute to broad, truncate apex with more than 6 teeth;

hyaline cells with up to 6 non-ringed pores on convex surface with few or no pseudopores, 2–4 elliptic ringed pores on concave surface in corners or along commissures, internal commissural walls minutely papillose (best viewed in oblique sections), rarely smooth; chlorophyllous cells narrowly triangular in transverse section, more broadly exposed on convex surface, enclosed on concave surface.

Sexual condition

dioicous.

monoicous.

Capsule

with abundant pseudostomata on surface of capsule.

Spores

33–40 µm.

31–43 µm; coarsely papillose on both proximal and distal surfaces, raised Y-mark sculpture on distal surface;

proximal laesura moderately long, 0.4–0.7 spore radius.

Branch

fascicles with 2 spreading and 1–2 pendent branches.;

branch stems green but sometimes reddish at proximal end, with cortex enlarged with conspicuous retort cells.

fascicles with 2 short-spreading and 3 long-tapering pendent branches.

Sphagnum majus

Sphagnum strictum

Phenology Capsules common, mature early to mid summer.
Habitat Pioneer species among grasses on peaty sand, pine barrens, burned-over savannas, seeps in mountainous areas inland
Elevation low to high elevations
Distribution
North America; Eurasia
from FNA
AL; FL; GA; LA; MD; ME; NC; NJ; SC; VA; NB; NF; NS; Europe
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Though they seldom if ever overlap ecologically, Sphagnum strictum and S. squarrosum both usually have squarrose branch leaves, but S. squarrosum has a lingulate fringed stem leaf that differs greatly from the triangular and entire-margined stem leaf of S. strictum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branch leaf hyaline cells on convex leaf surface with 1-2 pores per fibril interval, these usually less than 1/3 the cell diameter.
subsp. majus
1. Branch leaf hyaline cells on convex leaf surface with 1 pore per fibril interval, these more than 1/3 the cell diameter.
subsp. norvegicum
Source FNA vol. 27, p. 70. FNA vol. 27, p. 56.
Parent taxa Sphagnaceae > Sphagnum > sect. Cuspidata Sphagnaceae > Sphagnum > sect. Rigida
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Subordinate taxa
S. majus subsp. majus, S. majus subsp. norvegicum
Synonyms S. cuspidatum var. majus S. compactum var. expositum, S. garberi, S. mexicanum
Name authority (Russow) C. E.O. Jensen: in Botaniske Forening København, Festskrift, 106. (1890) Sullivant: Musc. Allegh., 201. (1846)
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