Sphagnum lescurii |
Sphagnum steerei |
|
---|---|---|
Lescur's sphagnum |
steere's sphagnum |
|
Habit | Plants moderate-sized to robust; upright, prostrate, or aquatic; green, pale yellow, golden brown, dark brown, tinged with red in exposed sites and purplish in aquatic forms; capitulum rounded and often strongly twisted. | Plants moderate-sized to large, compact and stiff-stemmed with upswept branches; dark green and brown, dark chesnut brown, brown to blackish brown, often with bluish tinge when dry; forming dense low to moderately tall hummocks. |
Stem(s) | leaves lingulate to ovate-lingulate, 1.3–2 mm; apex truncate to rounded, usually denticulate; hyaline cells typically fibrillose for 1/2 of leaf or more, often 1–2-septate, convex surface with 4–12 or more pores per cell along the commissures, concave surface with fewer pores. |
leaves to 1.2 × 0.8 mm; rarely hemiisophyllous; hyaline cells mostly nonseptate, comb-lamellae usually absent, but sometimes weak. |
Branches | usually curving, often large and tumid. |
short and blunt-tipped, leaves spreading. |
Branch leaves | broadly ovate to ovate-lanceolate, 1.3–2.5 mm, greatly elongated in aquatic forms, straight or infrequently subsecund or subsquarrose; hyaline cells with 10–22 pores along the commissures on the convex surface, no or fewer pores per cell (1–8) on the concave surface. |
ovate to ovate-elliptical, 1.6–2.2 × 1 mm; hyaline cells on proximal half of convex surface with round to elliptic pores along the commissures; comb-lamellae on hyaline cell wall where overlying chlorophyllous cells; chlorophyllous cells broadly triangular in transverse section and well-enclosed on convex surface, end wall not thickened. |
Sexual condition | dioicous. |
dioicous. |
Capsule | exserted, with few pseudostomata. |
unknown. |
Spores | 27–34 µm; finely papillose on both surfaces, with distinct raised Y-mark sculpture (indistinctly bifurcated Y-mark) on the distal surface; proximal laesura less than 0.5 spore radius. |
unknown. |
Branch | fascicles with 2(rarely 3) spreading and 1–2(–3) pendent branches. |
fascicles with 2 spreading and 1 pendent branch.; branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the end walls of cortical cells, cortical cell walls usually with large round pores. |
Sphagnum lescurii |
Sphagnum steerei |
|
Habitat | Weakly minerotrophic in a broad range of wetlands, often of an aquatic or periodically dried character | Commonly in firm-bottomed poor to medium fen vegetation, subarctic to arctic regions |
Elevation | low to moderate elevations | low to moderate elevations |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NF; NS; Europe
|
AK; MB; NT; NU; QC; YT; e Asia |
Discussion | Sporophytes are uncommon in Sphagnum lescurii, which may be the most phenotypically variable of all the North American Sphagnum species, and quite probably deserves some taxonomic splitting. The tremendous phenotypic plasticity of this species, however, makes it quite difficult to sort out the genotypic component of variability, and thus most sphagnologists since Warnstorf have avoided the temptation of splitting and have instead treated this as one very variable species. This is the approach maintained in this treatment. We have also chosen not to use the earlier name S. denticulatum because its type is a phenotypic morphotype not clearly assignable to the current concept of either S. auriculatum or S. lescurii (K. I. Flatberg, pers. comm.). Some of the American material assignable to S. lescurii is quite likely the same as the European species S. auriculatum, but much of our material is certainly not the same. Until more definitive data are available, we have chosen to continue to use the name S. lescurii. The large stem leaf will generally distinguish this from similar species of sect. Subsecunda. See also discussion under 55. S. inundatum and 61. S. platyphyllum. The names Sphagnum alabamae Warnstorf, S. aquatile Warnstorf, S. obesum (Wilson) Warnstorf, S. rufescens (Nees & Hornschuch) Warnstorf, and S. turgidulum Warnstorf also have been applied to this taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sphagnum steerei is usually easily recognized in the field by its dense growth habit and dark brown color. Its ecology is somewhat unclear due to taxonomic confusion with S. imbricatum in the strict sense and S. austinii (R. E. Andrus 1987). The latter is the only species in the imbricatum complex that occurs where S. steerei does. Contrary to the view of H. A. Crum (1997), S. steerei is quite distinct from S. affine and S. austinii. There is no range overlap with either of those species and the macroscopic appearance is also quite different. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 81. | FNA vol. 27, p. 54. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. orlandense, S. plicatum, S. wieboldtii | S. imbricatum var. arcticum |
Name authority | Sullivant: in A. Gray, Manual ed. 2, 611. (1856) | R. E. Andrus: Bryologist 90: 218. (1987) |
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