Sphagnum lescurii |
Sphagnum squarrosum |
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Lescur's sphagnum |
rough peat moss, shaggy peat, sphagnum, spiky bog-moss, spread-leaf peat moss, spreading-leaf bog moss |
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Habit | Plants moderate-sized to robust; upright, prostrate, or aquatic; green, pale yellow, golden brown, dark brown, tinged with red in exposed sites and purplish in aquatic forms; capitulum rounded and often strongly twisted. | Plants robust, stiff; green, pale green, yellow-green; large terminal bud; typically as loose carpets in coniferous forests. |
Stem(s) | leaves lingulate to ovate-lingulate, 1.3–2 mm; apex truncate to rounded, usually denticulate; hyaline cells typically fibrillose for 1/2 of leaf or more, often 1–2-septate, convex surface with 4–12 or more pores per cell along the commissures, concave surface with fewer pores. |
green to red-brown; 2–3 superficial cortical layers.; stem leaves shorter than branch leaves, ovate-lingulate to oblong-lingulate, 1.6–1.8 × 1–1.2 mm; hyaline cells mostly nonseptate. |
Branches | usually curving, often large and tumid. |
long and tapering with distinct squarrose spreading leaves, often terete in tundra forms. |
Branch leaves | broadly ovate to ovate-lanceolate, 1.3–2.5 mm, greatly elongated in aquatic forms, straight or infrequently subsecund or subsquarrose; hyaline cells with 10–22 pores along the commissures on the convex surface, no or fewer pores per cell (1–8) on the concave surface. |
larger than stem leaves, 1.9–2.8 mm, conspicuously squarrose from ovate-hastate base and abruptly narrowed 1/2–1/3 distance from apex into involute-concave acumen, often terete in tundra forms; hyaline cells convex on both surfaces, non-ringed pores at ends and corners of cells, ringed pores on concave surface (4–8/cell) and nonringed pores (2–4/cell) on convex surface, internal commissural walls smooth or indistinctly papillose, chlorophyllous cells ovate triangular with widest part at or close to the convex surface. |
Sexual condition | dioicous. |
monoicous. |
Capsule | exserted, with few pseudostomata. |
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Spores | 27–34 µm; finely papillose on both surfaces, with distinct raised Y-mark sculpture (indistinctly bifurcated Y-mark) on the distal surface; proximal laesura less than 0.5 spore radius. |
17–30 µm; proximal surface finely papillose, distal surface smooth with raised bifurcated Y-mark sculpture; proximal laesura more than 0.5 spore radius. |
Branch | fascicles with 2(rarely 3) spreading and 1–2(–3) pendent branches. |
fascicles with 2 spreading and 2–3 pendent branches.; branch stems with 1–2 layers of cortical cells. |
Sphagnum lescurii |
Sphagnum squarrosum |
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Phenology | Sporophytes abundant, capsules mature early to mid summer. | |
Habitat | Weakly minerotrophic in a broad range of wetlands, often of an aquatic or periodically dried character | Forming loose carpets in rich habitats such as wet coniferous forests, Thuja swamps, karrs, medium fens, and stream margins |
Elevation | low to moderate elevations | low to high elevations |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NF; NS; Europe
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AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; ME; MI; MN; MT; NC; NH; NJ; NY; OR; PA; SD; TN; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia; Pacific Islands (New Zealand)
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Discussion | Sporophytes are uncommon in Sphagnum lescurii, which may be the most phenotypically variable of all the North American Sphagnum species, and quite probably deserves some taxonomic splitting. The tremendous phenotypic plasticity of this species, however, makes it quite difficult to sort out the genotypic component of variability, and thus most sphagnologists since Warnstorf have avoided the temptation of splitting and have instead treated this as one very variable species. This is the approach maintained in this treatment. We have also chosen not to use the earlier name S. denticulatum because its type is a phenotypic morphotype not clearly assignable to the current concept of either S. auriculatum or S. lescurii (K. I. Flatberg, pers. comm.). Some of the American material assignable to S. lescurii is quite likely the same as the European species S. auriculatum, but much of our material is certainly not the same. Until more definitive data are available, we have chosen to continue to use the name S. lescurii. The large stem leaf will generally distinguish this from similar species of sect. Subsecunda. See also discussion under 55. S. inundatum and 61. S. platyphyllum. The names Sphagnum alabamae Warnstorf, S. aquatile Warnstorf, S. obesum (Wilson) Warnstorf, S. rufescens (Nees & Hornschuch) Warnstorf, and S. turgidulum Warnstorf also have been applied to this taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In its typical robust form with strongly squarrose branch leaves, Sphagnum squarrosum is unmistakeable. Smaller forms such as occur in the higher mountains may be difficult to identify accurately without careful examination of microscopic details. In the tundra there sometimes occur large, terete forms of S. squarrosum but these are usually considerably more robust than S. teres. See also discussion under 14. S. strictum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 81. | FNA vol. 27, p. 59. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. orlandense, S. plicatum, S. wieboldtii | S. squarrosum var. imbricatum |
Name authority | Sullivant: in A. Gray, Manual ed. 2, 611. (1856) | Crome: Samml. Deut. Laubm., 24. (1803) |
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