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junghuhn's sphagnum

fat peat-moss, papillose sphagnum

Habit Plants moderate-sized, soft, loosely tufted, slender, capitulum flat-topped to rounded; pale, dirty green, yellowish to brownish; without metallic lustre when dry. Plants moderate-sized to fairly robust; strong-stemmed and generally compact, capitulum usually not much enlarged; greenish brown to deep golden brown; forming compact carpets in floating mats and depressions as well as dense stands on hummock sides and low hummocks.
Stem(s)

leaves triangular-lingulate, 1.2–1.6 mm, broadly apex acute to narrowly truncate and toothed, border narrow or indistinct at base (less than 0.25 the width);

hyaline cells rhomboid, mostly 0–1-septate;

convex surface with membrane pleats, concave surface with 1–3 rounded membrane gaps occupying most of cell.

leaves to 1.3 × 0.7 mm; rarely hemiisophyllous;

hyaline cells non-ornamented, mostly septate.

Branches

somewhat 5-ranked.

generally short and blunt, leaves spreading.

Branch leaves

ovate-lanceolate, 1.3–2 mm, strongly concave, apex strongly involute;

margins entire to somewhat toothed near apex, hyaline cells on convex surface with numerous ringed elliptic pores (6–10) along commissures, concave surface mostly aporose except near margins;

Sexual condition dioicous or monoicous.

broadly ovate, 1.7 × 1 mm;

hyaline cells on convex surface with round to elliptic pores along the commissures, hyaline cell walls covered with papillae where overlying chlorophyllous cells; chlorophyllous cells trapezoidal to truncate-elliptic in transverse section, equally exposed on both surfaces or less exposed on convex surface, end walls thickened.

Sexual condition

dioicous.

Capsule

with numerous pseudostomata.

Spores

21–23 µm; minutely papillose.

26–36 µm; more roughly papillose on distal surface than proximal surface, distinct raised, bifurcated-Y mark sculpture on distal surface;

proximal laesura 0.5 spore radius or more.

Branch

fascicles with 2 spreading and 1–2 pendent branches.

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the interior end walls, mostly with 1 pore per cell on superficial cell wall.

Sphagnum junghuhnianum

Sphagnum papillosum

Phenology Capsules mature mid to late summer.
Habitat Shady, seepy cliffs Very common in very poor to poor fen mire habitats where it is often a major peat former, but scarce to absent in truly ombrotrophic peatlands sites
Elevation low elevations low to moderate elevations
Distribution
from FNA
BC; e Asia
from FNA
AK; CA; CT; DE; IL; IN; MD; ME; MI; MN; NC; NH; NJ; NY; OH; OR; PA; SC; VT; WA; WI; WV; BC; NB; NF; NS; ON; PE; QC; YT; Eurasia
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Discussion

Sphagnum junghuhnianum in the flora area is known only from the Queen Charlotte Islands.

Sporophytes of Sphagnum junghuhnianum were not seen. Three other large, brown species of sect. Acutifolia have stem leaves without fimbriate to lacerate apices, S. subnitens (forms without red color), S. subfulvum, and S. flavicomans. Sphagnum flavicomans has a more pointed stem leaf and a darker brown color as well as a strongly different ecology and range. Both S. subnitens and S. subfulvum have a glossy sheen when dry that is lacking in S. junghuhnianum. Sexual condition and spore characters were taken from H. A. Crum (1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphagnum papillosum is often easily field-identifiable by its rich golden brown to dark brown color and short, blunt branches. Nearly all specimens have the papillae on the branch leaf chorophyll cells but a few smooth forms have been found. Such forms will have stem leaves with divided hyaline cells whereas in the confusable species S. palustre and S. centrale such cells are rare or absent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 95. FNA vol. 27, p. 53.
Parent taxa Sphagnaceae > Sphagnum > sect. Acutifolia Sphagnaceae > Sphagnum > sect. Sphagnum
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. junghuhnianum subsp. pseudomolle, S. pseudomolle S. cymbifolium var. papillosum, S. papillosum var. laeve, S. papillosum var. sublaeve, S. waghornei
Name authority Dozy & Molkenboer: Verh. Kon. Akad. Wetensch., Afd. Natuurk. 2: 8. (1854) Lindberg: Acta Soc. Sci. Fenn. 10: 280. (1872)
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