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contorted sphagnum, twisted peat-moss

Habit Plants moderate-sized and moderately weak-stemmed to moderately stiff; green, brownish green to brown; capitulum flat-topped and 5-radiate, terminal bud often visible. Plants moderate to small-sized, weak-stemmed to spawling; green, yellow-green to golden brown; capitulum usually large and flat with curved branches; green, yellow-green, or golden brown.
Stem(s)

leaves triangular to lingulate-triangular, equal to or more than 0.8 mm, spreading to appressed;

apex acute to apiculate, hyaline cells mostly efibrillose and nonseptate.

leaves triangular-lingulate to lingulate, 0.7–1.4 mm;

apex rounded-obtuse and weakly denticulate;

hyaline cells nonseptate, mostly efibrillose, and, if porose, with more pores per cell on the concave surface (3–6) than on the convex surface (0–2).

Branches

± straight and somewhat tapered, usually 5-ranked, leaves not greatly elongated at branch distal end.

somewhat curved, leaves spreading.

Branch leaves

narrowly ovate-lanceolate, greater than 1.2 mm, straight, slightly undulate and weakly recurved when dry, margins entire;

hyaline cells on convex surface with 1 pore per cell in apical end, on concave surface with round wall thinnings in the cell ends and angles; chlorophyllous cells in transverse section triangular to ovate-triangular and well-enclosed on the concave surface.

ovate to ovate-lanceolate, 1.2–2 mm;

subsecund;

hyaline cells with numerous tiny pores in a continuous line along the commissures on the convex surface, no or scattered pores on the concave surface.

Sexual condition

dioicous.

dioicous.

Capsule

exserted, with scattered pseudo-stomata.

Spores

24–33 µm; finely papillose on the superficial surface.

22–28 µm;

papillose on both surfaces;

proximal laesurae more than 0.5 spore radius.

Branch

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems green and often reddish at proximal end, with cortex enlarged with conspicuous retort cells.

fascicles with 2–3 spreading and 2–3(4) pendent branches.

Sphagnum isoviitae

Sphagnum contortum

Habitat Forming carpets in a wide variety of poor to medium fen habitats of both mire edge and mire wide character, not found in ombrotrophic mires Very minerotrophic, sometimes found in slightly basic mires, intolerant of shade
Elevation low to moderate elevations low to moderate elevations
Distribution
from FNA
CT; IN; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; VA; VT; WV; AB; NF; NS; QC; Europe
from FNA
AK; CA; CO; CT; ID; IL; IN; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; OR; PA; RI; VT; WA; WI; WV; AB; BC; NF; NS; ON; QC; Eurasia
Discussion

Sporophytes are uncommon in Sphagnum isoviitae. See discussion under 26. S. brevifolium and 28. S. fallax for distinction from these similar species. Sphagnum isoviitae has no range overlap with S. pacificum, the other North American species of the S. recurvum complex with apiculate stem leaves; the sharply recurved branch leaves of the latter, however, would separate it easily in any case. Spore features are those given by Flatberg.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sporophytes are rare in Sphagnum contortum. This species is often associated with S. warnstorfii, S. centrale, Campylium stellatum, and Calliergonella cuspidata. The relatively small size, curved capitulum branches, and loosely spreading, subsecund branch leaves separate this species along with S. subsecundum. See also discussion under 61. S. platyphyllum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 68. FNA vol. 27, p. 80.
Parent taxa Sphagnaceae > Sphagnum > sect. Cuspidata Sphagnaceae > Sphagnum > sect. Subsecunda
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Name authority Flatberg: J. Bryol. 17: 2, figs. 1, 2. (1992) Schultz: Prodr. Fl. Starg. Suppl., 64. (1819)
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