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common brown peat-moss, rusty bogmoss, rusty peat moss, sphagnum

steere's sphagnum

Habit Plants small and slender, stiff and usually compact, capitulum small and flat-topped; typically deep reddish brown, also greenish brown in shaded habitats and in early seasonal growth, without metallic lustre when dry. Plants moderate-sized to large, compact and stiff-stemmed with upswept branches; dark green and brown, dark chesnut brown, brown to blackish brown, often with bluish tinge when dry; forming dense low to moderately tall hummocks.
Stem(s)

leaves lingulate, 0.8–1.3 mm;

apex broadly rounded and entire to lacerate, sometimes slightly mucronate or slightly denticulate;

hyaline cells rhombic, 0–1(–2)-septate, usually efibrillose.

leaves to 1.2 × 0.8 mm; rarely hemiisophyllous;

hyaline cells mostly nonseptate, comb-lamellae usually absent, but sometimes weak.

Branches

long and slender to short and compact, unranked to 5-ranked.

short and blunt-tipped, leaves spreading.

Branch leaves

ovate-lanceolate, 1.1–1.3 mm, straight, concave, apex strongly involute;

margins entire, hyaline cells on convex surface with round to elliptic pores along the commissures, grading from small pores near the leaf apex to large pores near the base, concave surface with large round pores in proximal marginal regions of leaf.

ovate to ovate-elliptical, 1.6–2.2 × 1 mm;

hyaline cells on proximal half of convex surface with round to elliptic pores along the commissures;

comb-lamellae on hyaline cell wall where overlying chlorophyllous cells; chlorophyllous cells broadly triangular in transverse section and well-enclosed on convex surface, end wall not thickened.

Sexual condition

dioicous.

dioicous.

Capsule

unknown.

Spores

17–30 µm, finely papillose on proximal surface and pusticulate on distal surface;

proximal laesura less than 0.5 spore radius.

unknown.

Branch

fascicles with 2 spreading and 1–2 pendent branches.

fascicles with 2 spreading and 1 pendent branch.;

branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the end walls of cortical cells, cortical cell walls usually with large round pores.

Sphagnum fuscum

Sphagnum steerei

Phenology Capsules mature late summer.
Habitat Mires, hummocks, fens Commonly in firm-bottomed poor to medium fen vegetation, subarctic to arctic regions
Elevation low to high elevations low to moderate elevations
Distribution
from FNA
AK; CA; CO; CT; ID; IL; IN; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NY; OH; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia
[WildflowerSearch map]
from FNA
AK; MB; NT; NU; QC; YT; e Asia
Discussion

Sphagnum fuscum is common in ombrotrophic mires and alpine mountain summits where it may form small to large hummocks to 1 m in height, more infrequently in weakly minerotrophic mires and richer fens.

Sporophytes are common in Sphagnum fuscum, which is associated with S. angustifolium, S. fallax, S. magellanicum, S. papillosum, and more infrequently with S. teres, and S. warnstorfii in richer sites. Very widespread but generally easily recognized, it is the only small brown hummock-forming species of sect. Acutifolia over most of its range. There are some significant variations in this species. The stem leaves can vary from having a rounded, entire apex to having a somewhat flat and lacerate apex. The branches also vary from being unranked and slender to 5-ranked and blunt. The color also can vary from a light to a dark brown. There does not seem, however, to be any consistent pattern to these variations and thus no taxonomic recognition has been given to them. See also discussion under 73. S. flavicomans.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphagnum steerei is usually easily recognized in the field by its dense growth habit and dark brown color. Its ecology is somewhat unclear due to taxonomic confusion with S. imbricatum in the strict sense and S. austinii (R. E. Andrus 1987). The latter is the only species in the imbricatum complex that occurs where S. steerei does. Contrary to the view of H. A. Crum (1997), S. steerei is quite distinct from S. affine and S. austinii. There is no range overlap with either of those species and the macroscopic appearance is also quite different.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 94. FNA vol. 27, p. 54.
Parent taxa Sphagnaceae > Sphagnum > sect. Acutifolia Sphagnaceae > Sphagnum > sect. Sphagnum
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Synonyms S. acutifolium var. fuscum, S. tenuifolium, S. vancouveriense S. imbricatum var. arcticum
Name authority (Schimper) H. Klinggraff: Schriften Phys.-Ökon. Ges. Konigsberg 13: 4. (1872) R. E. Andrus: Bryologist 90: 218. (1987)
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