Sphagnum fuscum |
Sphagnum mendocinum |
|
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common brown peat-moss, rusty bogmoss, rusty peat moss, sphagnum |
Mendocino sphagnum |
|
Habit | Plants small and slender, stiff and usually compact, capitulum small and flat-topped; typically deep reddish brown, also greenish brown in shaded habitats and in early seasonal growth, without metallic lustre when dry. | Plants moderately robust and lax; terminal bud somewhat enlarged; yellow-green to light brownish green. |
Stem(s) | leaves lingulate, 0.8–1.3 mm; apex broadly rounded and entire to lacerate, sometimes slightly mucronate or slightly denticulate; hyaline cells rhombic, 0–1(–2)-septate, usually efibrillose. |
leaves broadly oblong-triangular, 1.2–1.5 mm; mostly appressed to stem; apex obtuse; hyaline cells narrow, usually nonseptate, efibrillose and aporose on convex surface near apex, on concave surface usually efibrillose with irregular pores along commissures in distal portion of leaf. |
Branches | long and slender to short and compact, unranked to 5-ranked. |
with loosely imbricate leaves; often 5-ranked; leaves little to somewhat elongated at distal end. |
Branch leaves | ovate-lanceolate, 1.1–1.3 mm, straight, concave, apex strongly involute; margins entire, hyaline cells on convex surface with round to elliptic pores along the commissures, grading from small pores near the leaf apex to large pores near the base, concave surface with large round pores in proximal marginal regions of leaf. |
ovate, ovate-lanceolate to lanceolate; 2–3.5 mm; flat and undulate at margins and recurved at apex when dry; straight; margins entire; hyaline cells on concave surface with very numerous, small ringed or unringed pores along the commissures, 5–12 in distal portion of leaf and 20–27 in proximal portion, convex surface with 5–15 pores per cell in distal portion of leaf and 14–21 in proximal portion, pores usually without a ring; chlorophyllous cells triangular to trapezoidal in transverse section and exposed slightly on concave surface. |
Sexual condition | dioicous. |
dioicous. |
Spores | 17–30 µm, finely papillose on proximal surface and pusticulate on distal surface; proximal laesura less than 0.5 spore radius. |
ca. 30 µm; very slightly roughened. |
Branch | fascicles with 2 spreading and 1–2 pendent branches. |
fascicles with 2 spreading and 2–3 pendent branches.; branch stems green, with cortical cells in 1 layer with conspicuous necks. |
Sphagnum fuscum |
Sphagnum mendocinum |
|
Phenology | Capsules mature late summer. | |
Habitat | Mires, hummocks, fens | Submerged or floating in weakly minerotrophic wet depressions of alder swamps, coniferous swamps, Spiraea thickets, sedge fens, raised bogs, and drainage ditches in mires |
Elevation | low to high elevations | low to moderate elevations |
Distribution |
AK; CA; CO; CT; ID; IL; IN; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NY; OH; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Eurasia
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AK; CA; ID; MT; OR; WA; BC |
Discussion | Sphagnum fuscum is common in ombrotrophic mires and alpine mountain summits where it may form small to large hummocks to 1 m in height, more infrequently in weakly minerotrophic mires and richer fens. Sporophytes are common in Sphagnum fuscum, which is associated with S. angustifolium, S. fallax, S. magellanicum, S. papillosum, and more infrequently with S. teres, and S. warnstorfii in richer sites. Very widespread but generally easily recognized, it is the only small brown hummock-forming species of sect. Acutifolia over most of its range. There are some significant variations in this species. The stem leaves can vary from having a rounded, entire apex to having a somewhat flat and lacerate apex. The branches also vary from being unranked and slender to 5-ranked and blunt. The color also can vary from a light to a dark brown. There does not seem, however, to be any consistent pattern to these variations and thus no taxonomic recognition has been given to them. See also discussion under 73. S. flavicomans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sporophytes of Sphagnum mendocinum are uncommon. Although the unique branch leaf porosity makes it unmistakable microscopically, it may be confused with other species in the field. It is unusual ecologically in that it seems to overlap both the carpet- and lawn-forming habits—in other words it seems intermediate between S. recurvum and S. cuspidatum, in the broad sense. Among the species that it overlaps floristically, it is more robust and darker colored than S. pacificum. The latter also has an apiculate stem leaf compared to the more or less obtuse stem leaves of S. mendocinum. The more wet-growing S. majus var. majus and S. majus var. norvegicum, with which it slightly overlaps in habitat, have branch leaves that are strongly elongated at the distal branch ends whereas those of S. mendocinum are not. See also discussion under 40. S. obtusum. Sexual condition and spore characters are taken from from H. A. Crum (1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 94. | FNA vol. 27, p. 72. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. acutifolium var. fuscum, S. tenuifolium, S. vancouveriense | S. mendocinum var. gracilescens, S. mendocinum var. recurvum, S. mendocinum var. robustum |
Name authority | (Schimper) H. Klinggraff: Schriften Phys.-Ökon. Ges. Konigsberg 13: 4. (1872) | Sullivant: Icon. Musc., suppl.: 12. (1874) |
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