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fringe bogmoss, fringe peat-moss, sphagnum

Mendocino sphagnum

Habit Plants typically small and slender, larger and compact in the Arctic, capitulum small to moderate-sized, often with a conspicuous terminal bud; green, yellowish brown to brown; without metallic lustre when dry. Plants moderately robust and lax; terminal bud somewhat enlarged; yellow-green to light brownish green.
Stem(s)

leaves spatulate to broad-spatulate, 0.8–1.5(–2) mm, strongly lacerate across the broad apex and often part way down the margins, border scarcely to strongly broadened at base (0.25 width of base or less);

hyaline cells rhomboid, efibrillose and often 1–2-septate.

leaves broadly oblong-triangular, 1.2–1.5 mm; mostly appressed to stem;

apex obtuse;

hyaline cells narrow, usually nonseptate, efibrillose and aporose on convex surface near apex, on concave surface usually efibrillose with irregular pores along commissures in distal portion of leaf.

Branches

not 5-ranked, quite terete, long, and slender Branch fascicles with 1– 2 spreading and 1–2 pendent branches.

with loosely imbricate leaves; often 5-ranked;

leaves little to somewhat elongated at distal end.

Branch leaves

ovate to ovate-lanceolate; 1.1–1.5(–2) mm, slightly concave, straight;

apex involute;

margins entire;

hyaline cells on convex surface with numerous pores along the commissures grading from small pores near leaf apex to large pores at base, concave surface with large round pores at leaf apex and along margins.

ovate, ovate-lanceolate to lanceolate; 2–3.5 mm;

flat and undulate at margins and recurved at apex when dry; straight;

margins entire;

hyaline cells on concave surface with very numerous, small ringed or unringed pores along the commissures, 5–12 in distal portion of leaf and 20–27 in proximal portion, convex surface with 5–15 pores per cell in distal portion of leaf and 14–21 in proximal portion, pores usually without a ring; chlorophyllous cells triangular to trapezoidal in transverse section and exposed slightly on concave surface.

Sexual condition

often monoicous.

dioicous.

Spores

20–27 µm, finely papillose on both surfaces;

proximal laesura less than 0.5 spore radius.

ca. 30 µm;

very slightly roughened.

Branch

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems green, with cortical cells in 1 layer with conspicuous necks.

Sphagnum fimbriatum

Sphagnum mendocinum

Habitat Submerged or floating in weakly minerotrophic wet depressions of alder swamps, coniferous swamps, Spiraea thickets, sedge fens, raised bogs, and drainage ditches in mires
Elevation low to moderate elevations
Distribution
from USDA
North America; South America; Eurasia; Pacific Islands (New Zealand)
[WildflowerSearch map]
from FNA
AK; CA; ID; MT; OR; WA; BC
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sporophytes of Sphagnum mendocinum are uncommon. Although the unique branch leaf porosity makes it unmistakable microscopically, it may be confused with other species in the field. It is unusual ecologically in that it seems to overlap both the carpet- and lawn-forming habits—in other words it seems intermediate between S. recurvum and S. cuspidatum, in the broad sense. Among the species that it overlaps floristically, it is more robust and darker colored than S. pacificum. The latter also has an apiculate stem leaf compared to the more or less obtuse stem leaves of S. mendocinum. The more wet-growing S. majus var. majus and S. majus var. norvegicum, with which it slightly overlaps in habitat, have branch leaves that are strongly elongated at the distal branch ends whereas those of S. mendocinum are not. See also discussion under 40. S. obtusum. Sexual condition and spore characters are taken from from H. A. Crum (1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants small and slender, capitulum small and with a conspicuous terminal bud; stem leaves fimbriate down the sides and weakly to moderately bordered at the base
subsp. fimbriatum
1. Plants moderate-sized and compact, capitulum moderate sized and lacking a conspicuous terminal bud; stem leaves entire down the sides and strongly bordered at the base
subsp. concinnum
Source FNA vol. 27, p. 92. FNA vol. 27, p. 72.
Parent taxa Sphagnaceae > Sphagnum > sect. Acutifolia Sphagnaceae > Sphagnum > sect. Cuspidata
Sibling taxa
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. mendocinum, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
S. affine, S. alaskense, S. andersonianum, S. angermanicum, S. angustifolium, S. annulatum, S. aongstroemii, S. arcticum, S. atlanticum, S. austinii, S. balticum, S. bartlettianum, S. beothuk, S. bergianum, S. brevifolium, S. capillifolium, S. carolinianum, S. centrale, S. compactum, S. contortum, S. cribrosum, S. cuspidatum, S. cyclophyllum, S. fallax, S. fimbriatum, S. fitzgeraldii, S. flavicomans, S. flexuosum, S. fuscum, S. girgensohnii, S. henryense, S. imbricatum, S. inexspectatum, S. inundatum, S. isoviitae, S. jensenii, S. junghuhnianum, S. kenaiense, S. lenense, S. lescurii, S. lindbergii, S. macrophyllum, S. magellanicum, S. majus, S. mcqueenii, S. microcarpum, S. mirum, S. mississippiense, S. molle, S. obtusum, S. oregonense, S. orientale, S. pacificum, S. palustre, S. papillosum, S. perfoliatum, S. perichaetiale, S. platyphyllum, S. portoricense, S. pulchrum, S. pylaesii, S. quinquefarium, S. recurvum, S. riparium, S. rubellum, S. rubiginosum, S. rubroflexuosum, S. russowii, S. sitchense, S. splendens, S. squarrosum, S. steerei, S. strictum, S. subfulvum, S. subnitens, S. subsecundum, S. subtile, S. talbotianum, S. tenellum, S. tenerum, S. teres, S. torreyanum, S. trinitense, S. tundrae, S. viride, S. warnstorfii, S. wilfii, S. wulfianum
Subordinate taxa
S. fimbriatum subsp. concinnum, S. fimbriatum subsp. fimbriatum
Synonyms S. mendocinum var. gracilescens, S. mendocinum var. recurvum, S. mendocinum var. robustum
Name authority Wilson & Hooker: in J. D. Hooker, Fl. Antarct., 398. (1847) Sullivant: Icon. Musc., suppl.: 12. (1874)
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