Flora of North America species comparison

leaves triangular-ovate, more than 1.2 mm, usually appressed;

apex acute to apiculate, hyaline cells rarely septate or porose, apical region often fibrillose.

leaves may be less fibrillose or efibrillose and less porose or aporose than the branch leaves, often septate, a distinct border of narrow linear chlorophyllous cells often along margins and at base, and with a greater width:length ratio than branch leaves in anisophyllous forms, partly differentiated in hemiisophyllous forms, and identical in isophyllous forms.

mostly unranked to weakly 5-ranked, often conspicuously falcate, leaves greatly elongated at distal end.

typically dimorphic as spreading and pendent branches, but some species lack branches or branches are not clearly differentiated, pendent branches typically more slender than spreading branches and with a tendency to adhere to and cover the stem.

ovate-lanceolate to lanceolate, 1.6–5 mm, falcate toward branch tips, when dry often undulate and recurved, rarely weakly serrulate along the margins in submerged forms, leaves from middle of spreading branches with length to width ratio less than or equal to 1:0.28;

hyaline cells length to width ratio in apical convex surface region 8:1 or more, convex surface with 0–1 small round pores at apex, concave surface with faint round wall thinnings in cell apices and angles; chlorophyllous cells triangular to trapezoidal in transverse section, broadly exposed on the convex surface and exposed slightly on the concave surface.

with 2/5 phyllotaxy, of a 1-stratose network of alternating chlorophyllous and hyaline cells;

hyaline cells usually S-shaped, rarely rhomboid, nearly always strengthened with conspicuous spiral fibrils, small to large, round to elliptic and sometimes ringed pores occur along commissures or rarely on cell lumen, convex surface typically with more pores per cell than concave surface; chlorophyllous cells may be enclosed on both surfaces, more broadly exposed on one surface or equally exposed on both surfaces as viewed in transverse section, adjacent cell walls typically smooth, but various types of cell wall projections may be clearly visible in transverse section.

dioicous.

dioicous or monoicous; stalked globose antheridia borne at the tips of branches usually with swollen colored tips of branches near capitulum; long-necked archegonia borne on short branches singly surrounded by perichaetial leaves that are typically longer than branch leaves.

spherical, brown to black, lacking an annulus or peristome with a operculum convex;

spore sac amphithecial in origin, over-arching columella.

membranous.

29–38 µm; covered with large papillae on both surfaces, appearing pusticulate;

proximal laesura less than 0.5 spore radius.

tetrahedral, with prominent trilete mark, fine to coarse superficial surface, distal surface may have raised Y-mark, bifurcated Y-mark sculpture, or none.

fascicles with 2 spreading and 2–3 pendent branches.;

branch stems green, but often pinkish at the proximal ends, with cortex enlarged with conspicuous retort cells.

fascicles typically with 2 spreading and 1–2 pendent branches, but there may be up to 12(–14) per fascicle.;

branch stems typically green, with a superficial layer of inflated retort cells; these grouped or solitary, usually porose at the distal end with a conspicuous or inconspicuous neck.

typically 1-stratose, gametophyte developing from lateral margin.