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hairy sand-spurrey, hairy sand-spurry, villous sand spurry

purple sand spurry, red sand-spurrey, red sand-spurry, spergulaire rouge

Habit Plants strongly perennial with branched, woody base, stout, 11–30 cm, stipitate-glandular in inflorescence or throughout. Plants annual or short-lived perennial, delicate, 4–25 cm, stipitate-glandular in inflorescence.
Taproots

becoming stout, woody.

slender to ± stout.

Stems

erect to ascending, often arcuately so, much-branched proximally;

main stem 0.4–1.3 mm diam. proximally.

erect to ascending or prostrate, usually much-branched proximally;

main stem 0.3–0.5 mm diam. proximally.

Leaves

stipules ± conspicuous, dull white, broadly lance-acuminate, 3–8 mm, apex mucronate;

blade filiform to linear, 1–4.2 cm, somewhat fleshy, apex apiculate or spine-tipped;

axillary leaves 2–4 per cluster.

stipules conspicuous, shiny white, lanceolate, 3.5–5 mm, apex long-acuminate;

blade filiform to linear, 0.4–1.5 cm, scarcely fleshy, apex apiculate to spine-tipped;

axillary leaves 2–4+ per cluster.

Pedicels

spreading to reflexed in fruit.

ascending to reflexed.

Flowers

sepals connate 0.5–0.7 mm proximally, lobes 1- or 3-veined, lance-ovate to lanceolate, 2.5–4 mm, to 5 mm in fruit, margins 0.1–0.6 mm wide, apex acute to acuminate but often briefly rounded at tip;

petals white, ± elliptic, 0.7–0.8 times as long as sepal;

stamens 7–10;

styles 0.4–0.6 mm.

sepals connate 0.5–0.7 mm proximally, lobes often 3-veined, lanceolate, (2–)2.5–3.2 mm, to 4 mm in fruit, margins 0.1–0.3 mm wide, apex obtuse to acute;

petals pink, obovate to ovate, 0.9–1 times as long as sepals;

stamens 6–10;

styles 0.6–0.8 mm.

Capsules

greenish to tan, (4–)5–6.5 mm, 1.1–1.3 times as long as sepals.

greenish to tan, 3.5–5 mm, 1–1.2 times as long as sepals.

Seeds

reddish brown to dark brown, often with submarginal groove, broadly ovate, plump, 0.4–0.5 mm, smooth, often sculptured with parallel, wavy lines, papillae often present;

wing often present, white, 0.1–0.2 mm wide, margins irregular.

red-brown to dark brown, with submarginal groove, broadly ovate or ± truncate, angular at broad end, plump, 0.4–0.6 mm, sculpturing of parallel, wavy lines, margins with peglike papillae (30x);

wing absent.

Cymes

simple to 3-compound.

simple to 3+-compound or flowers solitary and axillary.

2n

= 18, 27, 36, 54 (all Europe).

Spergularia villosa

Spergularia rubra

Phenology Flowering spring. Flowering spring–fall.
Habitat Sandy slopes and bluffs, clay ridges and plains, disturbed areas Open forests, gravelly glades, meadows, mud flats, roadsides, disturbed places
Elevation 0-500 m (0-1600 ft) 0-2400 m (0-7900 ft)
Distribution
from FNA
CA; OR; Mexico (Baja California); South America (Chile?) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; CO; CT; DC; DE; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; UT; VA; VT; WA; WI; WY; BC; NB; NF; NS; ON; PE; QC; YT; SPM; Europe; Asia [Introduced in North America; introduced in South America, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Spergularia rubra was collected in 1901 on ballast in Alabama (Mohr, DS), the only record in the southeastern United States. It is the most widely distributed Spergularia species found outside of saline areas in the flora and has been in North America since at least the 1860s.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 22. FNA vol. 5, p. 19.
Parent taxa Caryophyllaceae > subfam. Polycarpoideae > Spergularia Caryophyllaceae > subfam. Polycarpoideae > Spergularia
Sibling taxa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. macrotheca, S. media, S. platensis, S. rubra, S. salina
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. macrotheca, S. media, S. platensis, S. salina, S. villosa
Synonyms Spergula villosa Arenaria rubra, Tissa rubra
Name authority (Persoon) Cambessèdes: in A. St.-Hilaire et al., Fl. Bras. Merid. 2: 129. (1830) (Linnaeus) J. Presl & C. Presl: Fl. ech. 94. 1819 
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