Spergularia salina |
Spergularia bocconi |
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lesser sea-spurrey, salt marsh sand-spurry, salt sandspurry, salt-marsh sand-spurrey, spergulaire des arais salés |
Boccone's sand spurry, Boccone's sand-spurrey, Boccone's sea-spurrey, Bocconi's sand-spurrey |
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Habit | Plants annual, delicate, 8–25(–30) cm, stipitate-glandular, at least in inflorescence. | Plants annual, ± delicate, 6–25+ cm, often densely stipitate-glandular, at least in inflorescence. |
Taproots | ± slender. |
± filiform. |
Stems | erect to ascending or prostrate, usually much-branched proximally; main stem 0.6–2(–3) mm diam. proximally. |
erect to spreading or sprawling, usually much-branched proximally; main stem 0.5–1 mm diam. proximally. |
Leaves | stipules inconspicuous, dull white, broadly triangular, 1.2–3.5 mm, longer than wide, apex acute to short-acuminate; blade linear, (0.8–)1.5–4 cm, fleshy, apex blunt to apiculate; axillary leaf clusters usually absent. |
stipules usually inconspicuous, dull white to tan, broadly triangular, 1.5–4.5 mm, apex acute to short-acuminate; blade ± linear, 1–4.2 cm, at least moderately fleshy, apex apiculate to spine-tipped; axillary leaves absent or 1–2 per cluster. |
Pedicels | reflexed and oriented to 1 side in fruit. |
often oriented to 1 side in fruit. |
Flowers | sepals connate 0.5–1 mm proximally, lobes often 3-veined, ovate to elliptic, 2.5–4.5 mm, to 4.8 mm in fruit, margins 0.1–0.5 mm wide, apex acute to rounded; petals white or pink to rosy, ovate to elliptic-oblong, 0.8–1 times as long as sepals; stamens (1–)2–3(–5); styles 0.4–0.7 mm. |
sepals connate 0.3–0.6 mm proximally, lobes often 3-veined, ovate to elliptic-oblong, 2.2–3.5 mm, to 4.5 mm in fruit, margins 0.2–0.5 mm wide, apex acute to rounded; petals white or pink to rosy, ovate to obovate, 0.8–1 times as long as sepals; stamens 8–10; styles 0.4–0.6 mm. |
Capsules | greenish to tan, 2.8–6.4 mm, 1–1.5 times as long as sepals. |
greenish, 3(–4) mm, 1–1.2 times as long as sepals. |
Seeds | light brown to reddish brown, with submarginal groove, broadly ovate, ± plump, 0.5–0.7(–0.8) mm, dull, ± smooth, often with gland-tipped papillae (30x); wing usually absent or incomplete. |
light brown, with submarginal groove, broadly ovate, plump, 0.4–0.6 mm, somewhat shiny, smooth to minutely roughened, margins with peglike papillae (40x); wing absent. |
Cymes | simple to 3+-compound or flowers solitary and axillary. |
simple to 6+-compound. |
2n | = 18? (Asia), 36 (Europe). |
= 18? (Africa), 36 (Europe). |
Spergularia salina |
Spergularia bocconi |
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Phenology | Flowering summer–early fall. | Flowering spring. |
Habitat | Mud flats, alkaline fields, sandy river bottoms, sandy coasts, salt marshes, saline highway edges (Great Lakes region) | Salt marshes, alkaline places, sandy soils |
Elevation | 0-1400 m (0-4600 ft) | 0-400 m (0-1300 ft) |
Distribution |
AL; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; KY; LA; MA; MD; ME; MI; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; TX; UT; VA; WA; WI; WY; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK; SPM; Eurasia; almost cosmopolitan via introduction [Introduced and native]
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CA; OR; sw Europe (Mediterranean region) [Introduced in North America]
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Discussion | While Spergularia salina may be native in coastal areas and some inland saline sites in much of the cited range, populations in the Great Lakes region are introduced where, as in S. media, highway and sidewalk salt runoff has created favorable habitats. Variety tenuis has been distinguished from var. salina by some authors as follows: cyme crowded versus lax, sepals 1.6–3.8 mm versus 2.4–5 mm, mature capsules 3–4.4 mm versus 3.6–6.4 mm, respectively. Due to the extreme overlap in morphologic features as well as geographic ranges, var. tenuis is not recognized here. The name Spergularia marina var. leiosperma (Kindberg) Gurke has been applied to plants with smooth seeds but, as pointed out by R. P. Rossbach (1940), separation of plants with smooth versus papillose seeds is not practical. Some authors believe that the correct name for this species is Spergularia marina. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The spelling of the epithet bocconi, often “corrected” to bocconii, is debatable. It commemorates Paolo Boccone, suggesting a correction to bocconei, but he also used the Latinized form Bocconus, allowing bocconi. We have used bocconi, following the first usage by Scheele. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 22. | FNA vol. 5, p. 20. |
Parent taxa | Caryophyllaceae > subfam. Polycarpoideae > Spergularia | Caryophyllaceae > subfam. Polycarpoideae > Spergularia |
Sibling taxa | ||
Synonyms | S. marina, S. marina var. tenuis, S. salina var. tenuis, S. tenuis, Tissa marina | Alsine bocconi |
Name authority | J. Presl & C. Presl: Fl. ech. 95. (1819) | (Scheele) Graebner: in P. F. A. Ascherson et al., Syn. Mitteleur. Fl. 5(1): 849. (1919) |
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