The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

beach sand-spurrey, beach sand-spurry, sticky sand spurry, sticky sand-spurrey

purple sand spurry, red sand-spurrey, red sand-spurry, spergulaire rouge

Habit Plants strongly perennial with branched, woody base, stout, 5–40 cm, densely stipitate-glandular in inflorescence or throughout. Plants annual or short-lived perennial, delicate, 4–25 cm, stipitate-glandular in inflorescence.
Taproots

becoming stout, woody.

slender to ± stout.

Stems

erect or ascending to prostrate, usually branched proximally;

main stem 0.8–3 mm diam. proximally.

erect to ascending or prostrate, usually much-branched proximally;

main stem 0.3–0.5 mm diam. proximally.

Leaves

stipules conspicuous, dull white to tan, narrowly triangular, 4.5–11 mm, apex long-acuminate;

blade linear, (0.6–)1–5.5 cm, fleshy, apex apiculate to spine-tipped;

axillary leaves 1–2+ per cluster.

stipules conspicuous, shiny white, lanceolate, 3.5–5 mm, apex long-acuminate;

blade filiform to linear, 0.4–1.5 cm, scarcely fleshy, apex apiculate to spine-tipped;

axillary leaves 2–4+ per cluster.

Pedicels

erect, divergent, or reflexed in fruit.

ascending to reflexed.

Flowers

sepals connate 0.5–1.8 mm proximally, lobes often 3-veined, ovate to lanceolate, 4.5–7 mm, to 8 mm in fruit, margins 0.3–0.7 mm wide, apex blunt to rounded;

petals white or pink to rosy, elliptic to obovate, 0.9–1.3 times as long as sepals in flower;

stamens 9–10;

styles 0.6–3 mm.

sepals connate 0.5–0.7 mm proximally, lobes often 3-veined, lanceolate, (2–)2.5–3.2 mm, to 4 mm in fruit, margins 0.1–0.3 mm wide, apex obtuse to acute;

petals pink, obovate to ovate, 0.9–1 times as long as sepals;

stamens 6–10;

styles 0.6–0.8 mm.

Capsules

tan, 4.6–10 mm, 0.8–1.4 times as long as sepals.

greenish to tan, 3.5–5 mm, 1–1.2 times as long as sepals.

Seeds

± red-brown, often with submarginal groove or depression, suborbiculate to pyriform, compressed, (0.6–)0.7–1.2 mm, smooth to faintly or prominently tuberculate or sculpturing of parallel, wavy lines or of low, rounded mounds, not papillate;

wing absent or white to reddish brown proximally, 0.1–0.3 mm wide, margins irregular.

red-brown to dark brown, with submarginal groove, broadly ovate or ± truncate, angular at broad end, plump, 0.4–0.6 mm, sculpturing of parallel, wavy lines, margins with peglike papillae (30x);

wing absent.

Cymes

simple to 3-compound or flowers solitary and axillary.

simple to 3+-compound or flowers solitary and axillary.

2n

= 18, 27, 36, 54 (all Europe).

Spergularia macrotheca

Spergularia rubra

Phenology Flowering spring–fall.
Habitat Open forests, gravelly glades, meadows, mud flats, roadsides, disturbed places
Elevation 0-2400 m (0-7900 ft)
Distribution
from FNA
CA; OR; WA; BC; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; CO; CT; DC; DE; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; UT; VA; VT; WA; WI; WY; BC; NB; NF; NS; ON; PE; QC; YT; SPM; Europe; Asia [Introduced in North America; introduced in South America, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Spergularia rubra was collected in 1901 on ballast in Alabama (Mohr, DS), the only record in the southeastern United States. It is the most widely distributed Spergularia species found outside of saline areas in the flora and has been in North America since at least the 1860s.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pink to rosy; styles 0.6-1.2 mm
var. macrotheca
1. Petals white; styles 1.2-3 mm
→ 2
2. Capsules 1.2-1.4 times as long as sepals; styles 1.2-1.9 mm
var. leucantha
2. Capsules 0.8-1 times as long as sepals; styles 2-3 mm
var. longistyla
Source FNA vol. 5, p. 18. FNA vol. 5, p. 19.
Parent taxa Caryophyllaceae > subfam. Polycarpoideae > Spergularia Caryophyllaceae > subfam. Polycarpoideae > Spergularia
Sibling taxa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. media, S. platensis, S. rubra, S. salina, S. villosa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. macrotheca, S. media, S. platensis, S. salina, S. villosa
Subordinate taxa
S. macrotheca var. leucantha, S. macrotheca var. longistyla, S. macrotheca var. macrotheca
Synonyms Arenaria macrotheca Arenaria rubra, Tissa rubra
Name authority (Hornemann ex Chamisso & Schlechtendal) Heynhold: Alph. Aufz. Gew. 689. (1846) (Linnaeus) J. Presl & C. Presl: Fl. ech. 94. 1819 
Web links